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Taxonomy of the genus Chersobius Biodiversity of the family Testudinidae

Bibliography of the genus
Chersobius (Five-toed Dwarf Tortoises)

(Reptilia: Testudines: Testudinidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Chersobius in general

Hofmeyr, M.D.; Branch, W.R. 2018. The padloper’s tortuous path (Chelonia: Testudinidae): Two genera, not one. African Journal of Herpetology 67(2): 99-112.

Chersobius boulengeri

Boycott, R.C. 1989. Homopus boulengeri Karoo padloper; red padloper; Biltong tortoise (English). Karooskilpadjie; Rooiskilpadjie; Donderweerskilpad; Biltongskilpad (Afrikaans). Occasional Papers of the IUCN Species Survival Commission 5: 78-79.

Eglis, A. 1963. Nesting of a parrot-beaked tortoise. Herpetologica 19: 66-68.

Hewitt, J. 1931. Descriptions of some African tortoises. Annals of the Natal Museum 6(3): 461-505.

Rowlands, B.W. 1988. Sympatry in land tortoises - resource partitioning: a need for research. Naturalist (Port Elizabeth) 32(1): 16-21.

Wyk, J.C.P. van; Bates, M.F. 1999. Geographical distribution: Homopus boulengeri Duerden, 1906. Karroo Padloper. South Africa: Free State. African Herp News 29: 42-43.

Chersobius signatus

Bayoff, N. 1995. Observations and morphometric data on the Namaqualand speckled tortoise, Homopus signatus signatus (Gmelin, 1789), in South Africa. Chelonian Conservation and Biology 1(3): 215-220.

Bour, R. 1988. Taxonomic and nomenclatural status of Homopus signatus (Gmelin, 1789): Reptilia-Chelonii. Journal of the Herpetological Association of Africa 35: 1-6.

Boycott, R. 1986. A review of Homopus signatus (Schöpff) with notes on related species (Cryptodira: Testudinidae). Journal of the Herpetological Association of Africa 32: 10-16.

Branch, W.R.; Bauer, A.M.; Jackman, T.R.; Marais, J. 2007. Homopus signatus subsp. Speckled Padloper. African Herp News 43: 26-27.

Daniels, S.R.; Hofmeyr, M.D.; Henen, B.T.; Baard, E.H.W. 2010. Systematics and phylogeography of a threatened tortoise, the speckled padloper. Animal Conservation 13(3): 237-246.

Dobiey, M. 2006. Gesägte Flachschildkröte Homopus signatus signatus (Gmelin, 1789). Reptilia (D) 11(61): 51-54.

Hewitt, J. 1931. Descriptions of some African tortoises. Annals of the Natal Museum 6(3): 461-505.

Hofmeyr, M.D.; Henen, B.T.; Loehr, V.J.T. 2005. Overcoming environmental and morphological constraints: egg size and pelvic kinesis in the smallest tortoise, Homopus signatus. Canadian Journal of Zoology 83(10): 1343-1352.

Klerks, M. 2002. Adapting the Namaqualand Speckled Padloper; Homopus signatus signatus, to captive conditions. Turtle and Tortoise Newsletter 6: 30-32.

Kooijman, M. 2015. Constipation in a specimen of the Speckled Cape Tortoise, Homopus signatus (Gmelin, 1789). Radiata (English edition) 24(4): 16-24.

Kooijman, M. 2015. Darmverstopfung bei einem Exemplar der Gesägten Flachschildkröte, Homopus signatus (Gmelin, 1789). Radiata 24(4): 16-24.

Licitra, J.T. 2001. Homopus signatus signatus. Speckled Padloper. Maximum size. African Herp News 32: 14-15.

Licitra, J.T. 2001. Homopus signatus signatus. Speckled Padloper. Observations on natural diet. African Herp News 32: 12-13.

Loehr, V. 1999. Dietary requirements of captive hatchling Namaqualand Speckled Padlopers (Homopus s. signatus). African Herp News 28: 23-26.

Loehr, V. 1999. Husbandry, behavior, and captive breeding of the Namaqualand speckled padloper (Homopus signatus signatus). Chelonian Conservation and Biology 3(3): 468-473.

Loehr, V. 1999. Photoperiod, temperatures and breeding in captive Namaqualand Speckled Padlopers, Homopus s. signatus. African Herp News 28: 27-28.

Loehr, V.; Harris, T.J. 1999. Life history notes: Homopus signatus. Speckled Padloper. Natural diet. African Herp News 29: 33-34.

Loehr, V.J.T. 2001. Homopus signatus signatus. Namaqualand Speckled Padloper. Female mating activity. African Herp News 32: 17-18.

Loehr, V.J.T. 2002. Diet of the Namaqualand Speckled Padloper, Homopus signatus signatus, in early spring. African Journal of Herpetology 51(1): 47-55.

Loehr, V.J.T. 2002. Male aggression in captive Namaqualand Speckled Padlopers, Homopus signatus signatus. Bulletin of the Chicago Herpetological Society 37(1): 1.

Loehr, V.J.T. 2002. Population characteristics and activity patterns of the Namaqualand Speckled Padloper (Homopus signatus signatus) in the early spring. Journal of Herpetology 36(3): 378-389.

Loehr, V.J.T. 2003. Annual temperature fluctuation in the natural habitat of the Namaqualand Speckled Padloper, Homopus signatus signatus. Radiata 12(1): 25-27.

Loehr, V.J.T. 2004. A new thread-trailing method for small tortoises in densely structured habitats. Turtle and Tortoise Newsletter 7: 13-14.

Loehr, V.J.T. 2004. First recorded second generation breeding with the Namaqualand Speckled Padloper, Homopus signatus signatus. Radiata 13(1): 11-12.

Loehr, V.J.T. 2004. Growth of the Namaqualand Speckled Padloper, Homopus signatus signatus (Reptilia: Testudinidae). African Zoology 39(2): 309-313.

Loehr, V.J.T. 2004. Homopus signatus signatus. Namaqualand Speckled Padloper. Aggressive courtship behaviour. African Herp News 37: 18-19.

Loehr, V.J.T. 2005. Preliminary results of a long-term field study on the Namaqualand Speckled Padloper, Homopus signatus signatus. Emys 12(5): 4-17.

Loehr, V.J.T. 2006. Egg and hatchling characteristics of the Namaqualand Speckled Padloper (Homopus signatus signatus): preliminary data from a captive population. Chelonii 4: 273-275.

Loehr, V.J.T. 2006. Natural diet of the Namaqualand Speckled Padloper (Homopus signatus signatus). Chelonian Conservation and Biology 5(1): 149-152.

Loehr, V.J.T. 2008. Homopus signatus signatus (Gmelin, 1789). Namaqualand Speckled Padloper. Natural oviposition and incubation. African Herp News 44: 9-10.

Loehr, V.J.T. 2010. Structure and dynamics of a Namaqualand Speckled Tortoise (Homopus signatus signatus) population over 5 years of rainfall variation. Chelonian Conservation and Biology 9(2): 223-230.

Loehr, V.J.T. 2012. High body temperatures in an arid, winter-rainfall environment: thermal biology of the smallest tortoise. Journal of Arid Environments 82: 123-129.

Loehr, V.J.T. 2015. Small vernal home ranges in the Namaqualand Speckled Tortoise, Homopus signatus. Journal of Herpetology 49(3): 447-451.

Loehr, V.J.T. 2016. Wide variation in carapacial scute patterns in a natural population of Speckled Tortoises, Homopus signatus. African Journal of Herpetology 65(1): 47-54.

Loehr, V.J.T.; Henen, B.T.; Hofmeyr, M.D. 2004. Reproduction of the smallest tortoise, the namaqualand Speckled Padloper, Homopus signatus signatus. Herpetologica 60(4): 444-454.

Loehr, V.J.T.; Henen, B.T.; Hofmeyr, M.D. 2006. Shell characteristics and sexual dimorphism in the Namaqualand Speckled Padloper, Homopus signatus signatus. African Journal of Herpetology 55(1): 1-11.

Loehr, V.J.T.; Henen, B.T.; Hofmeyr, M.D. 2011. Reproductive responses to rainfall in the Namaqualand Speckled Tortoise. Copeia 2011(2): 278-284.

Loehr, V.J.T.; Hofmeyr, M.D.; Henen, B.T. 2007. Growing and shrinking in the smallest tortoise, Homopus signatus signatus: the importance of rain. Oecologia (Berlin) 153(2): 479-488.

Loehr, V.J.T.; Hofmeyr, M.D.; Henen, B.T. 2009. Small and sensitive to drought: consequences of aridification to the conservation of Homopus signatus signatus. African Journal of Herpetology 58(2): 116-125.

Loehr, V.J.T.; Schmalz, M. 2001. Homopus signatus cafer. Southern Speckled Padloper. Autumn activity. African Herp News 32: 16.

Loehr, V.J.T.; Stark, T.; Weterings, M.; Kuipers, H. 2015. Overcoming low environmental temperatures in the primary feeding season: low-level activity and long basking in the tortoise Homopus signatus. Amphibia-Reptilia 36(3): 207-214.

Philippen, H.D. 2008. Gesägte Flachschildkröte Homopus signatus signatus (Gmelin, 1789). Marginata 17: 31-34.

Saus, A. 2005. Beobachtungen zur Nahrungsaufnahme in Relation zur Luftfeuchtigkeit einer Nachzucht von Homopus signatus signatus. Schildkröten im Fokus 2(1): 29-32.

Schmidt, F.; Engel-Brecht, K. 2006. Natural history notes: Homopus signatus (Speckled Padloper). Maximum male size. Herpetological Review 37(2): 216.

Stark, T. 2013. Natural history notes: Homopus signatus (Gmelin, 1789). Diet. African Herp News 59: 19-21.

Chersobius solus

Branch, W.R. 1992. Homopus 'bergeri' - a wrong name for a new tortoise from southern Namibia. Journal of the Herpetological Association of Africa 40: 11.

Branch, W.R. 2007. A new species of tortoise of the genus Homopus (Chelonia: Testudinidae) from southern Namibia. African Journal of Herpetology 56(1): 1-21.

Karl, H.V.; Tichy, G. 1999. Die Taxonomie von Homopus bergeri Lindholm 1906 (Testudines: Testudinoidea). Mauritiana 17(2): 277-284.

Schleicher, A. 2004. Erfahrungen und Beobachtungen bei der Haltung der Namibia-Flachschildkröte, Homopus sp. ("Homopus bergeri"), im Freilandgehege in Namibia sowie Bericht über die erfolgreiche Nachzucht dieser Art. Radiata 13(4): 3-12.

Schleicher, A. 2004. Experiences with keeping and breeding the Namibian Cape Tortoise Homopus sp. ("Homopus bergeri"), in a Namibian outdoor enclosure. Radiata (English edition) 13(4): 3-12.

Schleicher, A.; Loehr, V.J.T. 2001. Husbandry, behavior, and captive breeding of the Nama padloper, Homopus bergeri, from southwestern Namibia. Chelonian Conservation and Biology 4(1): 165-170.

Schleicher, A.; Loehr, V.J.T. 2006. Husbandry, behaviour and captive breeding of the Nama Padloper, Homopus sp. pp. 459-466. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Siebenrock, F. 1909. Homopus bergeri Ldh., eine Testudo-Art aus der Geometrica-Gruppe. Zoologischer Anzeiger 34: 623-625.