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Taxonomy of the genus Conolophus Biodiversity of the family Iguanidae

Bibliography of the genus
Conolophus (Galapagos Land Iguanas)

(Reptilia: Sauria: Iguanidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Conolophus in general

Carpenter, C.C. 1969. Behavioral and ecological notes on the Galapagos land iguanas. Herpetologica 25: 155-164.

Cayot, L.; Snell, H.L.; Llerena, W.; Snell, H.M. 1994. Conservation biology of Galapagos reptiles: twenty-five years of successful research and management. pp. 297-305. In: Murphy, J.B.; Adler, K. & Collins, J.T. (eds.). Captive management and conservation of amphibians and reptiles. SSAR (Society for the Study of Amphibians and Reptiles). 408 pp. (Contributions to Herpetology no. 11).

Denburgh, J. van; Slevin, J.R. 1913. The Galapagoan lizards of the genus Tropidurus; with notes on the iguanas of the genera Conolophus and Amblyrhynchus. (Expedition of the California academy of Sciences to the Galapagos islands, 1905-1906, 9.). Proceedings of the California Academy of Sciences 2: 132-202.

Dowling, H.G. 1964. Goats and hawks - A new theory of predation on the land iguana. Animal Kingdom 67: 51-56.

Eibl-Eibesfeldt, I. 1984. The large iguanas of the Galapagos Islands. pp. 157-173. In: Perry, R. (ed.). Key environments. Galapagos. Pergamon Press, Oxford, New York etc. 321 pp.

Etheridge, R.E. 1982. Checklist of the iguanine and Malagasy iguanid lizards. pp. 7-37. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Hagen, W. von 1937. Galapagos land iguanas. Nature Magazine 29: 368-370.

Higgins, P.J. 1978. The Galapagos iguanas: models of reptilian differentiation. Bioscience 28(8): 512-515.

Higgins, P.J.; Rand, C.S.; Haynes, J.L. 1974. Galapagos iguanas: Amblyrhynchus and Conolophus serum protein relationships. Journal of Experimental Zoology 189(2): 255-260.

Luecker, H.; Feiler, A. 2002. Beobachtung einer Paarung zwischen Meerechsen-Männchen (Amblyrhynchus cristatus) und Landleguan-Weibchen (Conolophus subcristatus) auf Plaza-Sur/Galapagos-Inseln sowie Beobachtungen an einem adulten Gattungshybriden (Amblyrhynchus X Conolophus). Elaphe 10(4): 49-54.

Marquez, C.; Snell, H.; Snell, H.; Rea, S.; Wilson, M.; Cepeda, F. 1986. The ten year struggle to save the endangered land iguanas. Noticias de Galapagos 44: 9-11.

Rassmann, K.; Markmann, M.; Trillmich, F.; Tautz, D. 2004. Tracing the evolution of the Galapagos iguanas: a molecular approach. pp. 71-83. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Reynolds, R.P. 1982. Programa de herpetologia. Estacion Cientifica Charles Darwin Informe Anual 1981: 51-55.

Snell, H.; Snell, H. 1980. 1979 annual report, land iguana conservation program. Charles Darwin Research Station Annual Report 1979: 36-51.

Snell, H.; Snell, H. 1981. Land iguana conservation. Charles Darwin Research Station Annual Report 1980: 129-131.

Snell, H.L.; Christian, K.A. 1985. Energetics of Galapagos land iguanas: a comparison of two island populations. Herpetologica 41(4): 437-442.

Snell, H.L.; Snell, H.M.; Tracy, C.R. 1984. Variation among populations of Galapagos land iguanas (Conolophus): contrasts of phylogeny and ecology. Biological Journal of the Linnean Society 21(1-2): 185-207.

Switak, K.H. 2002. Landleguane der Galapagos-Inseln. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 55(1): 18-21.

Tzika, A.C.; Rosa, S.F.P.; Fabiani, A.; Snell, H.L.; Snell, H.M.; Marquez, C.; Tapia, W.; Rassmann, K.; Gentile, G.; Milinkovitch, M.C. 2008. Population genetics of Galapagos land iguana (genus Conolophus) remnant populations. Molecular Ecology 17(23): 4943-4952.

Vries, T. de 1974. Opmerkingen over taxonomie en ecologie van de reptielen van de Galapagos-eilanden. 2. Landleguanen en zeeleguanen. Lacerta 32(6): 95-106.

Wyles, J.S.; Sarich, V.M. 1983. Are the Galapagos iguanas older than the Galapagos? Molecular evolution and colonization models for the archipelago. pp. 177-186. In: Bowman, R.I.; Berson, M. & Leviton, A.E. (eds.). Patterns of evolution in Galapagos organisms. Pacific Division AAAS, San Francisco. 568 pp.

Conolophus marthae

Donegan, T.M. 2009. Type specimens, samples of live individuals and the Galapagos pink land iguana. Zootaxa 2201: 12-20.

Dubois, A. 2009. Endangered species and endangered knowledge. Zootaxa 2201: 26-29.

Frick, M.G. 2010. Misconceptions about the ability of researchers to relocate the holotype of the Galapagos pink land iguana through the use of a passive integrated transponder. Zootaxa 2694: 57-58.

Gentile, G.; Fabian, A.; Marquez, C.; Snell, H.L.; Snell, H.M.; Tapia, W.; Sbordoni, V. 2009. An overlooked pink species of land iguana in the Galapagos. Proceedings of the National Academy of Sciences of the United States of America 106(2): 507-511.

Gentile, G.; Snell, H. 2009. Conolophus marthae sp. nov. (Squamata, Iguanidae), a new species of land iguana from the Galapagos archipelago. Zootaxa 2201: 1-10.

Giambattista, L. di; Gentile, G. 2018. A molecular protocol to distinguish syntopic Galápagos Land Iguanas (Conolophus marthae and C. subcristatus) from faecal samples. Herpetology Notes 11: 97-100.

Nemesio, A. 2009. On the live holotype of the Galapagos pink land iguana, Conolophus marthae Gentile & Snell, 2009 (Squamata: Iguanidae): is it an acceptable exception? Zootaxa 2201: 21-25.

Conolophus pallidus

Christian, K. 1980. Sante Fe land iguana project. Charles Darwin Research Station Annual Report 1979: 146-147.

Christian, K.; Porter, W.P.; Tracy, C.R. 1986. Core areas within the home ranges of Galapagos land iguanas, Conolophus pallidus. Journal of Herpetology 20(2): 272-276.

Christian, K.; Tracy, C.R.; Porter, W.P. 1983. Seasonal shifts in body temperature and use of microhabitats by Galapagos land iguanas (Conolophus pallidus). Ecology (Washington, D.C.) 64(3): 463-468.

Christian, K.A. 1982. Relations among space, time and thermal niche axes of the Galapagos land iguana Conolophus pallidus. Dissertation Abstracts International B Sciences and Engineering 42(11): 4322.

Christian, K.A.; Clavijo, I.E.; Cordero-Lopez, N.; Elias-Maldonado, E.E.; Franco, M.A.; Lugo-Ramirez, M.V.; Marengo, M. 1986. Thermoregulation and energetics of a population of Cuban iguanas (Cyclura nubila) on Isla Magueyes, Puerto Rico. Copeia 1986(1): 65-69.

Christian, K.A.; Tracy, C.R. 1980. An update on the status of Isla Santa Fe since the eradication of the feral goats. Noticias de Galapagos 31: 16-17.

Christian, K.A.; Tracy, C.R. 1981. The effect of the thermal environment on the ability of hatchling Galapagos land iguanas to avoid predation during dispersal. Oecologia (Heidelberg) 49(2): 218-223.

Christian, K.A.; Tracy, C.R. 1982. Reproductive behavior of Galapagos land iguanas, Conolophus pallidus, on Isla Santa Fe, Galapagos. pp. 366-379. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Christian, K.A.; Tracy, C.R. 1984. Physiological and ecological consequences of sleeping-site selection by the Galapagos land iguana (Conolophus pallidus). Ecology (Washington, D.C.) 65(3): 752-758.

Christian, K.A.; Tracy, C.R. 1985. Physical and biotic determinants of space utilization by the Galapagos land iguana (Conolophus pallidus). Oecologia (Heidelberg) 66(1): 132-140.

Christian, K.A.; Tracy, C.R.; Porter, W.P. 1984. Diet, digestion and food preferences of Galapagos land iguanas. Herpetologica 40(2): 205-212.

Fritts, T.H.; Snell, H.L.; Cayot, L.; MacFarland, C.; Earsom, S.; Marquez, C.; Llerena, W.; Llerena, F. 2000. Progress and priorities in research for the conservation of reptiles. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique Entomologie 70(Supplement): 39-45.

Mackie, R.I.; Rycyk, M.; Ruemmler, R.L.; Aminov, R.I.; Wikelski, M. 2004. Biochemical and microbiological evidence for fermentative digestion in free-living land iguanas (Conolophus pallidus) and marine iguanas (Amblyrhynchus cristatus) on the Galapagos Archipelago. Physiological and Biochemical Zoology 77(1): 127-138.

Rassmann, K. 1997. Evolutionary age of the Galapagos iguanas predates the age of the present Galapagos islands. Molecular Phylogenetics and Evolution 7(2): 158-172.

Thaller, M.C.; Migliore, L.; Márquez, C.; Tapia, W.; Cedeno, V.; Rossolini, G.M.; Gentile, G. 2010. Tracking acquired antibiotic resistance in commensal bacteria of Galapagos land iguanas: no man, no resistance. PLoS ONE 5(2): e8989, 1-4.

Tracy, C.R.; Christian, K.A. 1985. Are marine iguana tails flattened? British Journal of Herpetology 6(12): 434-435.

Wilhoft, D.C. 1958. Observations on preferred body temperature and feeding habits of some selected tropical iguanas. Herpetologica 14: 161-164.

Conolophus subcristatus

Anonymous. 1978. First Galapagos land iguana hatched in captivity. IZN (International Zoo News) 25(7): 26-27.

Anonymous. 1978. Land iguanas saved by incubation in captivity. Smithsonian Institution Research Reports 22: 6.

Anonymous. 1987. Repatriation of captive-bred land iguanas. Noticias de Galapagos 45: 4-5.

Baur, G. 1896. Professor Cope's criticisms of my drawings of the squamosal region of Conolophus subcristatus. American Naturalist 30: 327-329.

Cayot, L. 1993. Terrestrial iguanas in Baltra: an update. Galapagos Bulletin 1993(Fall): 4.

Cayot, L.; Menoscal, R. 1992. Land iguanas return to Baltra. Noticias de Galapagos 51: 11-13.

Cayot, L.J. 2008. The restoration of giant tortoise and land iguana populations in Galapagos. Galapagos Research 65: 39-43.

Celleri, Y. 1983. Repatriation of land iguanas (Conolophus subcristatus) to Cartago Bay on Isabela Island. Estacion Cientifica Charles Darwin Informe Anual 1982: 138-140.

Costantini, D.; Filippis, S.P. de; Marquez, C.; Snell, H.L.; Snell, H.M.; Tapia, W.; Brambilla, G.; Gentile, G. 2009. Temporal and spatial covariation of gender and oxidative stress in the Galapagos land iguana Conolophus subcristatus. Physiological and Biochemical Zoology 82(5): 430-437.

Dieckmann, M. 2014. [Species profile Conolophus subcristatus (GRAY, 1831)]. (In German). Iguana Rundschreiben 27(2): 16-20.

Ellis, H.I.; Ross, J.P. 1978. Field observations of cooling rates of Galapagos land iguanas (Conolophus subcristatus). Comparative Biochemistry and Physiology A Comparative Physiology 59(2): 205-209.

Fabiani, A.; Trucchi, E.; Rosa, S.; Márquez, C.; Snell, H.L.; Snell, H.M.; Aguilera, W.T.; Gentile, G. 2011. Conservation of Galapagos land iguanas: genetic monitoring and predictions of a long-term program on the island of Santa Cruz. Animal Conservation 14(4): 419-429.

Fritts, T.H.; Snell, H.L.; Cayot, L.; MacFarland, C.; Earsom, S.; Marquez, C.; Llerena, W.; Llerena, F. 2000. Progress and priorities in research for the conservation of reptiles. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique Entomologie 70(Supplement): 39-45.

Giambattista, L. di; Gentile, G. 2018. A molecular protocol to distinguish syntopic Galápagos Land Iguanas (Conolophus marthae and C. subcristatus) from faecal samples. Herpetology Notes 11: 97-100.

Harper, G.A.; Zabala, J.; Carrion, V. 2011. Monitoring of a population of Galapagos land iguanas (Conolophus subcristatus) during a rat eradication using brodifacoum. Occasional Papers of the IUCN Species Survival Commission 42: 309-312.

Hillaby, J. 1964. The riddle of the elderly iguanas. New Scientist 22: 438-439.

Hofkin, B.V.; Wright, A.; Altenbach, J.; Rassmann, K.; Snell, H.M.; Miller, R.D.; Stone, A.C.; Snell, H.L. 2003. Ancient DNA gives green light to Galapagos land iguana repatriation. Conservation Genetics 4(1): 105-108.

Marquez, C. 1986. Herpetology program of the CDRS. Estacion Cientifica Charles Darwin Informe Anual 1983: 28-29.

Oftedal, O.T.; Allen, M.E. 1996. Nutrition as a major facet of reptile conservation. Zoo Biology 15(5): 491-497.

Oliver, J.A. 1956. Lizards of the sea. Animal Kingdom 59: 148-151.

Olson, S.L. 2014. The early scientific history of Galapagos iguanas. Archives of Natural History 41(1): 141-153.

Rassmann, K. 1997. Evolutionary age of the Galapagos iguanas predates the age of the present Galapagos islands. Molecular Phylogenetics and Evolution 7(2): 158-172.

Rassmann, K.; Trillmich, F.; Tautz, D. 1997. Hybridization between the Galapagos land and marine iguana (Conolophus subcristatus and Amblyrhynchus cristatus) on Plaza Sur. Journal of Zoology (London) 242(4): 729-739.

Reynolds, R.P. 1981. Land iguanas (Conolophus subcristatus) on North Seymour Island. Noticias de Galapagos 34: 17-18.

Reynolds, R.P. 1982. Experimental repatriation of captive-reared land iguanas (Conolophus subcristatus) at Cartago Bay, Isabela. Noticias de Galapagos 36: 12-14.

Reynolds, R.P. 1983. Experimental repatriation of captive-reared land iguanas (Conolophus subcristatus) at Cartago Bay, Isabela. Ambio 12(3-4): 189.

Snell, H.; Snell, H. 1982. Rare land iguana breeds in captivity. Rephiberary 52: 3. (Reprinted from WWF Monthly Report, August 1981).

Snell, H.L. 1984. Evolutionary ecology of Galapagos land iguanas (Iguanidae: Conolophus). Dissertation Abstracts International B Sciences and Engineering 45(5): 1369.

Snell, H.L.; Snell, H.M.; Tracy, C.R. 1982. Reproductive ecology of the Plaza Sur land iguana population. Estacion Cientifica Charles Darwin Informe Anual 1981: 105-110.

Snell, H.L.; Tracy, C.R. 1985. Behavioral and morphological adaptations by Galapagos land iguanas (Conolophus subcristatus) to water and energy requirements of eggs and neonates. American Zoologist 25(4): 1009-1018.

Vries, T. de; Black, J. 1983. Of men, goats and guava - problems caused by introduced species in the Galapagos. Noticias de Galapagos 38: 18-21.

Wallner, L.M. 1995. The Charles Darwin Research Station: Darwins's legacy. Vivarium (Lakeside) 6(5): 26-29.

Werner, D.I. 1980. Galapagos land iguana, natural history and conservation survey. Charles Darwin Research Station Annual Report 1979: 142-145.

Werner, D.I. 1982. Social organization and ecology of land iguanas, Conolophus subcristatus, on Isla Fernandina, Galapagos. pp. 342-365. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Werner, D.I. 1983. Reproduction in the iguana Conolophus subcristatus on Fernandina Island, Galapagos: clutch size and migration costs. American Naturalist 121(6): 757-775.

Woram, J.M. 1992. That first iguana transfer. Noticias de Galapagos 51: 20-22.