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Taxonomy of the genus Cyclura Biodiversity of the family Iguanidae

Bibliography of the genus
Cyclura (Caribbean Ground Iguanas)

(Reptilia: Sauria: Iguanidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Cyclura in general

Alberts, A.; IUCN;SSC West Indian Iguana Specialist Group (ed.). 2000. West Indian Iguanas: Status Survey and Conservation Action Plan. International Union for the Conservation of Nature and Natural Resources (IUCN), Cambridge. 111 pp.

Alberts, A.C. 2004. Conservation strategies for West Indian rock iguanas (genus Cyclura): current efforts and future directions. Iguana (West Hartford) 11(4): 212-223.

Alberts, A.C. 2006. Kidnapping the Don Juans of Guantanamo. Conservation Behaviorist 4(1): 19-20.

Alberts, A.C. 2007. Behavioral considerations of headstarting as a conservation strategy for endangered Caribbean rock iguanas. Applied Animal Behaviour Science 102(3-4): 380-391.

Areste, M. 1998. Cyclura: the ground iguanas of the Caribbean. Reptilia (GB) 2: 52-57.

Barbour, T.; Noble, G.K. 1916. A revision of the lizards of the genus Cyclura. Bulletin of the Museum of Comparative Zoology 60: 139-164.

Cope, E.D. 1885. The large iguanas of the Greater Antilles. American Naturalist 19: 1005-1006.

Duval, J.J. 1983. Recommendations for the captive management of West Indian rock iguanas, (Cyclura). Annual Reptile Symposium on Captive Propagation and Husbandry Proceedings 6: 181-196.

Etheridge, R.E. 1982. Checklist of the iguanine and Malagasy iguanid lizards. pp. 7-37. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Garber, S. 1976. Biogeography and distribution of Iguana and Cyclura in the Greater Antilles. (Abstract). Herpetological Review 7(2): 83.

Grant, T.D.; Hudson, R.D. 2015. West Indian iguana Cyclura spp. reintroduction and recovery programmes: zoo support and involvement. pp. 49–55. In: Zoological Society of London (ed.). International Zoo Yearbook. Vol. 49. Reptile Conservation. Zoological Society of London, London. 442 pp.

Hayes, W.K.; Iverson, J.B.; Knapp, C.R.; Carter, R.L. 2012. Do invasive rodents impact endangered insular iguana populations. Biodiversity and Conservation 21(7): 1893-1899.

Hudson, R. 1995. The AZA Regional Collection Plan for large lizards: the need for space. American Zoo and Aquarium Association Regional Conference Proceedings 1995: 275-280.

Hudson, R. 1996. AZA Species Survival Plan profile: West Indian rock iguanas. Endangered Species Update 13(7-8): 9-10, 14.

Hudson, R.D.; Alberts, A.C. 2004. The role of zoos in the conservation of West Indian iguanas. pp. 274-289. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Iguana Specialist Group (formerly the West Indian Iguana Specialist Group). . www.iucn-isg.org/actionplan/ch2/rhinocerous.php

Knapp, C.R. 2004. Ecotourism and its potential impact on iguana conservation in the Caribbean. pp. 290-301. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Knapp, C.R.; Hudson, R.D. 2004. Translocation strategies as a conservation tool for West Indian iguanas: evaluations and recommendations. pp. 199-209. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Knapp, C.R.; Owens, A.K. 2004. Diurnal refugia and novel ecological attributes of the Bahamian boa, Epicrates striatus fowleri (Boidae). Caribbean Journal of Science 40(2): 265-270.

Köhler, G. 1997. Zur Eiablage von Grossleguanen in der Natur und im Terrarium. Iguana Rundschreiben 10(1)(17): 32-36.

Lemm, J.M.; Alberts, A.C. 2012. Cyclura: natural history, husbandry, and conservation of West Indian rock iguanas. Academic (Elsevier), London. 239 pp.

Malone, C.L.; Wheeler, T.; Taylor, J.F.; Davis, S.K. 2000. Phylogeography of the Caribbean rock iguana (Cyclura): implications for conservation and insights on the biogeographic history of the West Indies. Molecular Phylogenetics and Evolution 17(2): 269-279.

Martins, E.P.; Lamont, J. 1998. Estimating ancestral states of a communicative display: a comparative study of Cyclura rock iguanas. Animal Behaviour 55(6): 1685-1706.

Noegel, R.P. 1989. Husbandry of west Indian rock iguanas Cyclura at Life Fellowship Bird Sanctuary, Seffner. International Zoo Yearbook 28: 131-135.

Pasachnik, S.A.; Martin-Velez, V. 2017. An evaluation of the diet of Cyclura iguanas in the Dominican Republic. Herpetological Bulletin 140: 6-12.

Ruiter, M. de 1993. Hoffnung für den Jamaika-Leguan. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 46(6): 348.

Sanlley Castro, C.; Duval, J. 1979. Reproducción en cautividad de iguanas del género Cyclura. Zoodom 3(2): 13-18.

Schwartz, A.; Carey, M. 1977. Systematics and evolution in the West Indian iguanid genus Cyclura. Studies on the fauna of Curaçao and other Caribbean Islands 53(173): 15-97.

Townson, S. 1980. West Indian iguanas of the genus Cyclura: the threat of habitat destruction in the Cayman Islands. British Journal of Herpetology 6(3): 101-104.

Cyclura carinata

Alberts, A.C. 2001. Turks and Caicos iguana rescue and restoration program, Caribbean. Re-introduction News 20: 22.

Auffenberg, W. 1982. Feeding strategy of the Caicos ground iguana, Cyclura carinata. pp. 84-116. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Auth, D.L. 1980. The thermal biology of the Turks and Caicos Islands rock iguana Cyclura carinata. Dissertation Abstracts International B Sciences and Engineering 41(6): 2082.

Bissell, A.N.; Martins, E.P. 2004. Behavior and ecology of rock iguanas, II: population differences. pp. 109-118. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Bryan, J.J.; Gerber, G.P.; Welch, M.E.; Stephen, C.L. 2007. Re-evaluating the taxonomic status of the Booby Cay iguana, Cyclura carinata bartschi. Copeia 2007(3): 734-739.

Cochran, D.M. 1931. New Bahaman reptiles. Journal of the Washington Academy of Sciences 21(3): 39-41.

Gerber, G.P. 2007. Turks and Caicos iguana translocation program, Bahama Archipelago. Re-introduction News 26: 53-55.

Gerber, G.P.; Alberts, A. 2004. Turks and Caicos Iguana, Cyclura carinata carinata: 2003 project update. Iguana (West Hartford) 11(1): 37.

Iverson, J.B. 1978. The impact of feral cats and dogs on populations of the West Indian rock iguana, Cyclura carinata. Biological Conservation 14(1): 63-73.

Iverson, J.B. 1979. Behaviour and ecology of the rock iguana Cyclura carinata. Bulletin of the Florida State Museum Biological Sciences 24(3): 175-358.

Iverson, J.B. 1985. Lizards as seed dispensers? Journal of Herpetology 19(2): 292-293.

Martins, E.P.; Lacy, K.E. 2004. Behavior and ecology of rock iguanas, I: evidence for an appeasement display. pp. 101-108. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Mitchell, N. 2003. Another chance for the Turks and Caicos Rock Iguana. Iguana (West Hartford) 10(1): 10-14.

Mitchell, N.C.; Haeffner, R.; Fulford, M.; Clerveaux, W. 2000. Geographic distribution: Cyclura carinata carinata (Turks and Caicos Rock Iguana). Herpetological Review 31(4): 253-254.

Mitchell, N.C.; Haeffner, R.; Veer, V.; Fulford Gardner, M.; Clerveaux, W.; Veitch, C.R.; Mitchell, G. 2002. Cat eradication and the restoration of endangered iguanas (Cyclura carinata) on Long Cay, Caicos Bank, Turks and Caicos Islands, British West Indies. Occasional Papers of the IUCN Species Survival Commission 27: 206-212.

Norton, R.L.; Clarke, N.V. 1992. Notes on the rock iguana (Cyclura carinata) of the Caicos Islands. Florida Field Naturalist 20(2): 45-46.

Reading, R.P.; Haeffner, R.; Veer, V.; Mitchell, N.C. 2011. Re-introduction of Turks and Caicos rock iguana to Long Cay, Turks and Caicos Banks. pp. 120-124. In: Soorae, P.S. Global re-introduction perspectives, 2011: more case studies from around the globe. IUCN/SSC Re-introduction Specialist Group & Environment Agency, Abu Dhabi. 250p.

Smith, G.R. 1992. Return of Cyclura carinata to Pine Cay, Turks and Caicos Islands, BWI. Herpetological Review 23(1): 21-23.

Welch, M.E.; Gerber, G.P.; Davis, S.K. 2004. Genetic structure of the Turks and Caicos Rock Iguana and its implications for species conservation. pp. 58-70. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Cyclura caymanensis

Blair, R.D. 1983. Dragons of Cayman. Rock iguanas cling to their islands. Oceans 16(2): 31-33.

Carey, W.M. 1966. Observations on the ground iguana Cyclura macleayi caymanensis on Cayman Brac, British West Indies. Herpetologica 22: 265-268.

Cyclura collei

Alberts, A. 1993. The most endangered lizard in the world: the Jamaican iguana, Cyclura collei. Vivarium (Lakeside) 5(1): 12-14.

Alberts, A.C. 2007. Behavioral considerations of headstarting as a conservation strategy for endangered Caribbean rock iguanas. Applied Animal Behaviour Science 102(3-4): 380-391.

Baum, T. 2008. Jamaika, seine Leguane und der Riesenanolis Anolis garmani. Reptilia (D) 13(72): 73-80.

Gibson, R.C. 1993. A short study of captive Jamaican iguanas Cyclura collei at Hope Zoo, Jamaica. Dodo 29: 156-167.

Henningheim, E. 2013. [Health-screening and release of the Jamaican iguana]. (In Dutch). Lacerta 71(5): 208-216.

Hudson, R. 1994. Conservation strategy for the endangered Jamaican iguana: the role of zoos. American Zoo and Aquarium Association Regional Conference Proceedings 1994: 36-42.

Hudson, R. 1995. The AZA Regional Collection Plan for large lizards: the need for space. American Zoo and Aquarium Association Regional Conference Proceedings 1995: 275-280.

Hudson, R. 2001. The critically endangered Jamaican iguana of Hellshire Hills. Species 35: 6-7.

Kong, D. 1994. The Jamaican iguana. American Zoo and Aquarium Association Regional Conference Proceedings 1994: 107-108.

Lewis, D.S.; Vogel, P.; Wilson, B.S. 2008. Survival, dispersal, and home range of headstarted Jamaican iguanas, Cyclura collei. Caribbean Journal of Science 44(3): 423-425.

Mittermeier, R.A. 1972. Jamaica's endangered species. Oryx 11(4): 258-262.

Searcy, R.A.; Villers, L.M.; Reams, R.D.; Wyatt, J.E.; Pilarski, J. 2009. Captive reproduction of the Jamaican iguana (Cyclura collei). Zoo Biology 28(4): 343-349.

Tole, L. 2001. Habitat loss and anthropogenic disturbance in Jamaica's Hellshire Hills area. Biodiversity and Conservation 11(4): 575-598.

Veen, R. van; Wilson, B.S.; Grant, T.; Hudson, R. 2014. Where to now? An uncertain future for Jamaica's largest endemic vertebrate. Oryx 48(2): 169-170.

Vogel, P. 1994. Evidence of reproduction in a remnant population of the endangered Jamaican iguana, Cyclura collei (Lacertilia, Iguanidae). Caribbean Journal of Science 30(3-4): 234-241.

Vogel, P. 2004. Jamaican Iguana field research and recovery project: 2003 summary. Iguana (West Hartford) 11(1): 31-32.

Vogel, P.; Nelson, R.; Kerr, R. 1996. Conservation strategy for the Jamaican iguana, Cyclura collei. pp. 395-406. In: Powell, R. & Henderson, R.W. (eds.). Contributions to West Indian herpetology: a tribute to Albert Schwartz. SSAR (Society for the Study of Amphibians and Reptiles). 457 pp. (Contributions to Herpetology no. 12).

Welch, M.E.; Rasberry, A.; Grant, T.; Veen, R. van; Robinson, O.; Fleuchaus, D.; Wilson, B. 2012. Assessing the genetic impact of headstarting and potential for mitigation in the Jamaican Iguana, Cyclura collei. IRCF Reptiles & Amphibians 19(3): 228–229.

Wilson, B.S.; Alberts, A.C.; Graham, K.S.; Hudson, R.D.; Bjorkland, R.K.; Lewsi, D.S.; Lung, N.P.; Nelson, R.; Thompson, N.; Kunna, J.L.; Vogel, P. 2004. Survival and reproduction of repatriated Jamaican iguanas: headstarting as a viable conservation strategy. pp. 220-231. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Wilson, B.S.; Robinson, O.F.; Vogel, P. 2004. Status of the Jamaican iguana (Cyclura collei): assessing 15 Years of conservation effort. Iguana (West Hartford) 11(4): 224-231.

Woodley, J.D. 1980. Survival of the Jamaican iguana, Cyclura collei. Journal of Herpetology 14(1): 45-49.

Cyclura cornuta

Anonymous. 1978. Nacimiento eu Zoodom de Iguanas rinoceronte (Cyclura c. cornuta) de segunda generacion. Zoodom 3(1): 31-32.

Anonymous. 1991. Junge Nashornleguane im Exotarium geschlüpft. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 44(11): 690.

Anonymous. 1992. Animal collection. North of England Zoological Society Annual Report 1991: 21-31.

Anonymous. 1992. Zum 3. Mal Nachzucht der seltenen Nashornleguane im Exotarium. Herpetofauna (Weinstadt) 14(79): 25.

Banbury, B.L.; Ramos, Y.M. 2005. The rock iguanas of Parque Nacional Isla Cabritos. Iguana (West Hartford) 12(4): 256-261.

Baylan, T. 1984. Breeding the rhinoceros iguana Cyclura c. cornuta at Sydney Zoo. International Zoo Yearbook 23: 144-148.

Boylan, T. 1985. Captive management of a population of rhinoceros iguanas Cyclura cornuta cornuta, at Taronga Zoo, Sydney. pp. 491-494. In: Grigg, G.; Shine, R. & Ehmann, H. (eds.). Biology of Australian frogs and reptiles. Surrey Beatty & Sons Pty & the Royal Zoological Society of New South Wales, Chipping Norton (NSW). 527 pp.

Brice, S. 1995. A review of feeding and lighting requirements for captive herbivorous lizards: Cyclura and Iguana. Dodo 31: 120-139.

Carey, W.M. 1975. The rock iguana, Cyclura pinguis on Anegada, British Virgin Islands, with notes on Cyclura ricardi and C. cornuta. Bulletin of the Florida State Museum Biological Sciences 19(4): 189-233.

Friedrich, U. 1979. Das Echsenporträt. Der Nashorn-Leguan. Aquarien Magazin 13(9): 443.

Glor, R.E.; Powell, R.; Parmerlee, J.S. 2000. Cyclura cornuta. Catalogue of American Amphibians and Reptiles 709: 1-6.

Gross, R. 1992. Wissenswertes über den Nashornleguan, Cyclura cornuta. Iguana Rundschreiben 1992(1): 20-23.

Haast, W.E. 1969. Hatching rhinoceros iguanas Cyclura cornuta at the Miami Serpentarium. International Zoo Yearbook 9: 49.

Hartley, L.M.; Glor, R.E.; Sproston, A.L.; Powell, R.; Parmerlee, J.S. 2000. Germination rates of seeds consumed by two species of rock iguanas (Cyclura spp.) in the Dominican Republic. Caribbean Journal of Science 36(1-2): 149-151.

Hudson, R. 1995. The AZA Regional Collection Plan for large lizards: the need for space. American Zoo and Aquarium Association Regional Conference Proceedings 1995: 275-280.

Klingel, G.C. 1929. Lizard hunting in the Black Republic. Natural History (New York) 29(5): 451-464.

Müller-Mainz, L. 1908. Über das Gefangenleben des Nashornleguans (Metopocerus cornutus). Wochenschrift für Aquarien- und Terrarienkunde (Braunschweig) 5: 27-31.

Noble, G.K. 1923. Trailing the rhinoceros iguana. Journal Amer. Mus. Nat. Hist. New York 23: 541-558.

Oliver, J.A. 1956. Lizards of the sea. Animal Kingdom 59: 148-151.

Oschman, C. (ed.) 1993. [Rhinoceros iguana hatched after six months]. (In Afrikaans, English summary). Zoon (Pretoria) 1993(3): 9.

Ottenwalder, J.A. 1978. International zoo news. Santo Domingo, Dominican Republic. IZN (International Zoo News) 25(7): 45.

Ottenwalder, J.A. 1979. International zoo news. Santo Domingo, Dominican Republic. IZN (International Zoo News) 26(1): 41.

Peters, U. 1981. Drei bemerkenswerte Nachzuchten: Netzpython, Nashorn-Leguan und Gopher-Schildkröte. Aquarium (Bornheim) 15(139): 33-37.

Peters, U. 1982. The breeding of endangered reptiles, a success story. Zoologische Garten 52(1): 21-28.

Peters, U.W. 1991. Zaunleguane. Prächtige Kobolde aus den USA. Aquarium (Bornheim) 268: 47-48.

Petzold, H.G. 1967. Ein Nashornleguan, Cyclura cornuta mit Gabelschwanz im Tierpark Berlin. Aquarien Terrarien 14: 306-308.

Powell, R. 1999. Herpetology of Navassa Island, West Indies. Caribbean Journal of Science 35(1-2): 1-13.

Ramer, J. 2004. A survey of Ricord's Iguanas (Cyclura ricordii) and Rhinoceros Iguanas (Cyclura cornuta cornuta) in Isla Cabritos National Park, Dominican Republic 2003: a preliminary report. Iguana (West Hartford) 11(2): 88-97.

Ramer, J.C.; Maria, R.; Reichard, T.; Tolson, P.J.; Chen, T.C.; Holick, M.F. 2005. Vitamin D status of wild Ricord's iguanas (Cyclura ricordii) and captive and wild rhinoceros iguanas (Cyclura cornuta cornuta) in the Dominican Republic. Journal of Zoo and Wildlife Medicine 36(2): 188-191.

Sanlley Castro, C.; Duval, J. 1979. Reproducción en cautividad de iguanas del género Cyclura. Zoodom 3(2): 13-18.

Schmidt, J. 1997. Erfahrungen bei der Zucht von Cyclura-Arten. Iguana Rundschreiben 10(1)(17): 21-26.

Shaw, C.E. 1969. Breeding the rhinoceros iguana Cyclura cornuta cornuta at San Diego Zoo. International Zoo Yearbook 9: 45-48.

Tonge, S.; Bloxam, Q. 1984. Captive reproduction of rhinoceros iguanas (Cyclura c. cornuta) in indoor accommodation. Acta Zoologica et Pathologica Antverpiensia 78: 115-128.

Townson, S. 1977. Temperature requirements and thermoregulation of lizards - general comments with special reference to the rhinoceros iguana (Cyclura c. cornuta). Herptile 2(3): 42-46.

Vifias Roman, J.A. 1979. Ecological distribution of Dominican iguanas. IZN (International Zoo News) 26(3): 6-9.

Warkentin, T.; Kessner, R. 2011. Erfolgreiches Nachzucht-Jahr in der Reptilienabteilung des Zoo-Aquariums Berlin. Bongo 41: 51-56.

Wilke, H. 1981. Das Nashornleguan, Cyclura cornuta Bonnaterre-natürlicher Lebensraum und Zoohaltung. Naturwissenschaftlicher Verein Darmstadt Bericht N.F. 5: 8-11.

Wilke, H. 1981. Die Situation von Cyclura cornuta cornuta Bonnaterre in Zoologischen Garten, ihre natürlichen Lebensbedingungen und der Versuch einer Übertragung auf die Tiere des Zoo Frankfurt a. M. Zoologische Garten 51(3-4): 177-190.

Cyclura cychlura

Bailey, A.M. 1925. The iguanas of Bitter Guana Cay. Natural History (New York) 25(4): 390-393.

Banff, L.; Knapp, C.R.; Greene, D.U.; Buckner, S.D.; Iverson, J.B. 2012. Natural history notes: Cyclura cychlura figginsi (Exuma Island Rock Iguana). Rodent kill. Herpetological Review 43(3): 483.

Barbour, T. 1923. Another new Bahaman Iguana. Proceedings of the New England Zoological Club 8: 107-109.

Bosch, H.A.J. in den 1991. Wirtelschwanzleguane im Lobbecke-Museum Cyclura cychlura figginsi. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 44(1): 29-31.

Coenen, C. 1995. Observations on the Bahamian rock iguana of the Exumas. Bahamas Journal of Science 2(2): 8-14.

Colosimo, G.; Knapp, C.R.; Wallace, L.E.; Welch, M.E. 2014. Inferred vs realized patterns of gene flow: an analysis of population structure in the Andros Island Rock Iguana. PLoS ONE 9(9): e106963.

Hines, K.N. 2011. Effects of ecotourism on endangered Northern Bahamian Rock Iguanas (Cyclura cychlura). Herpetological Conservation and Biology 6(2): 250-259.

Hines, K.N. 2017. On the distribution of Northern Bahamian Rock Iguanas (Cyclura cychlura) in the Exuma Islands. Herpetological Review 48(1): 28-31.

Hines, K.N.; Iverson, J.B.; Valiulis, J.M. 2002. Natural history notes: Cyclura cychlura inornata (Allen Cays Rock Iguana). Bird predation. Herpetological Review 33(4): 306.

Iverson, J.B. 2004. Allen's Cays iguanas, Cyclura cychlura inornata: summary report for 2003. Iguana (West Hartford) 11(1): 29.

Iverson, J.B. 2007. Juvenile survival in the Allen Cays Rock Iguana (Cyclura cychlura inornata). Copeia 2007(3): 740-744.

Iverson, J.B.; Converse, S.J.; Smith, G.R.; Valiulis, J.M. 2006. Long-term trends in the demography of the Allen Cays Rock Iguana (Cyclura cychlura inornata): human disturbance and density-dependent effects. Biological Conservation 132(3): 300-310.

Iverson, J.B.; Francois, K.; Jollay, J.; Buckner, S.D.; Knapp, C.R. 2017. Natural history notes: Cyclura cychlura figginsi (Exuma Rock Iguana). Allogrooming. Herpetological Review 48(1): 188.

Iverson, J.B.; Hines, K.N.; Valiulis, J.M. 2004. The nesting ecology of the Allen Cays Rock Iguana, Cyclura cychlura inornata in the Bahamas. Herpetological Monographs 18: 1-36.

Iverson, J.B.; Knapp, C.R.; Hines, K.N.; Pasachnik, S.A.; Pieper, L. 2011. Natural history notes: Cyclura cychlura inornata (Allen Cays Rock iguana). Mortality. Herpetological Review 42(2): 274.

Iverson, J.B.; Mamula, M.R. 1989. Natural growth in the Bahaman iguana Cyclura cychlura. Copeia 1989(2): 502-505.

Iverson, J.B.; Pasachnik, S.A.; Knapp, C.R.; Buckner, S.D. 2006. Cyclura cychlura. Catalogue of American Amphibians and Reptiles 810: 1-8.

Iverson, J.B.; Smith, G.R.; Pieper, L. 2004. Factors affecting long-term growth of the Allen Cays Rock Iguana in the Bahamas. pp. 176-192. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Iverson, J.B.; Young, D.P.; Young, C.A.; Schofield, C. 2005. Natural history notes: Cyclura cychlura inornata (Allen Cays Iguana). Mortality. Herpetological Review 36(2): 175.

James, S.B.; Iverson, J.; Greco, V.; Raphael, B.L. 2006. Health assessment of Allen Cays Rock Iguana, Cyclura cychlura inornata. Journal of Herpetological Medicine and Surgery 16(3): 93-97.

Jungnickel, J. 2009. Cyclura cychlura inornata von Leaf Cay, Bahamas. Sauria (Berlin) 31(1): 39-40.

Knapp, C. 1995. A flora and fauna survey of Guana Cay, with an emphasis on its rock iguana. Bahamas Journal of Science 2(2): 2-7.

Knapp, C. 2004. 2003 research update for Exuma (Cyclura cychlura figginsi) and Andros (C. c. cychlura) Islands, Bahamas. Iguana (West Hartford) 11(1): 32-33.

Knapp, C.; Buckner, S. 1998. Geographic distribution: Cyclura cyclura figginsi (Exuma Island Ground Iguana). Herpetological Review 29(1): 51.

Knapp, C.; Buckner, S.; Feldman, A.; Roth, L. 1999. Status update and empirical field observations of the Andros rock iguana, Cyclura cychlura cychlura. Bahamas Journal of Science 7(1): 2-5.

Knapp, C.R. 2000. Home range and intraspecific interactions of a translocated iguana population (Cyclura cychlura inornata Barbour and Noble). Caribbean Journal of Science 36(3-4): 250-257.

Knapp, C.R. 2001. Status of a translocated Cyclura iguana colony in the Bahamas. Journal of Herpetology 35(2): 239-248.

Knapp, C.R. 2005. Working to save the Andros Iguana. Iguana (West Hartford) 12(1): 8-13.

Knapp, C.R. 2015. Staying high and dry: a unique nesting strategy for an insular iguana. Frontiers in Ecology and the Environment 13(2): 114-115.

Knapp, C.R.; Alvarez-Clare, S. 2016. Influence of morphological, chemical and physical leaf traits on food selection of a herbivorous iguana from the Bahamas. Journal of Tropical Ecology 32(1): 75-78.

Knapp, C.R.; Alvarez-Clare, S.; Pérez-Heydrich, C. 2010. The influence of landscape heterogeneity and dispersal on survival of neonate insular iguanas. Copeia 2010(1): 62-70.

Knapp, C.R.; Iverson, J.B.; Owens, A.K. 2006. Geographic variation in nesting behavior and reproductive biology of an insular iguana (Cyclura cychlura). Canadian Journal of Zoology 84(11): 1566-1575.

Knapp, C.R.; Malone, C.L. 2003. Patterns of reproductive success and genetic variability in a translocated iguana population. Herpetologica 59(2): 195-202.

Knapp, C.R.; Owens, A.K. 2005. Home range and habitat associations of a Bahamian iguana: implications for conservation. Animal Conservation 8(3): 269-278.

Knapp, C.R.; Owens, A.K. 2008. Natural history notes: Cyclura cychlura cychlura (Andros Iguana). Attemped predation. Herpetological Review 39(2): 221-222.

Knapp, C.R.; Owens, A.K. 2008. Nesting behavior and the use of termitaria by the Andros Iguana (Cyclura cychlura cychlura). Journal of Herpetology 42(1): 46-53.

Knapp, C.R.; Pagni, L. (eds.). 2011. Andros iguana: conservation action plan, 2005-2011. IUCN/SSC Iguana Specialist Group, Gland. 24 pp.

Owens, A.K.; Knapp, C.R. 2007. Natural history notes: Cyclura cychlura cychlura (Andros Iguana). Scoliosis; Kyphosis. Herpetological Review 38(4): 454-455.

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Smith, G.R.; Iverson, J.B. 2006. Changes in sex ratio over time in the endangered iguana Cyclura cychlura inornata. Canadian Journal of Zoology 84(10): 1522-1527.

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Cyclura lewisi

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Burton, F.J. 2005. Restoring a new wild population of Blue Iguanas (Cyclura lewisi) in the Salina Reserve, Grand Cayman. Iguana (West Hartford) 12(3): 166-174.

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Soifer, F.K. 1992. Conservation efforts on behalf of Cyclura nubila lewisii. Journal of Small Exotic Animal Medicine 1(3): 133-135.

Cyclura nubila

Alberts, A.C. 1995. Use of statistical models based on radiographic measurements to predict oviposition date and clutch size in rock iguanas (Cyclura nubila). Zoo Biology 14(6): 543-553.

Alberts, A.C. 1999. Developing recovery strategies for West Indian rock iguanas. Endangered Species Update 16(5): 107-110.

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Christian, K. 1987. Aspects of the life history of Cuban iguanas on Isla Magueyes, Puerto Rico. Caribbean Journal of Science 22(3-4): 159-164.

Christian, K.A. 1986. Physiological consequences of nighttime temperature for a tropical, herbivorous lizard (Cyclura nubila). Canadian Journal of Zoology 64(4): 836-840.

Christian, K.A.; Clavijo, I.E.; Cordero-Lopez, N.; Elias-Maldonado, E.E.; Franco, M.A.; Lugo-Ramirez, M.V.; Marengo, M. 1986. Thermoregulation and energetics of a population of Cuban iguanas (Cyclura nubila) on Isla Magueyes, Puerto Rico. Copeia 1986(1): 65-69.

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Fenske, R.; Grellmann, T. 1997. Beobachtungen an Kubaleguanen (Cyclura nubila Gray, 1831) auf der Insel Cayo Largo del Sur vor Kuba. Sauria (Berlin) 19(3): 3-7.

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Gonzalez-Rossell, A.; Berovides Alvarez, V.; Castaneira Colome, M.A. 2004. Variacion intrapoblacional de la densidad y las caracteristicas de los refugios de la iguana Cubana (Cyclura nubila nubila) en Cayo del Rosario, Archipielago de los Canarreos, Cuba. Revista Biologia (Havana) 18(1): 50-56.

Grellmann, T.; Fenske, R.; Helmdag, A. 1995. Beobachtungen der Kubaleguane Cyclura nubila Gray auf der Insel Cayo Largo del Sur vor Kuba. Sauria (Berlin) 17(3): 19-21.

Hardy, J.D. 1956. Notes on the Cuban iguana. Herpetologica 12: 323-324.

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Perera, A. 1985. Datos sobre abundancia y actividad de Cyclura nubila (Sauria: Iguanidae) en los alrededores de Cayo Largo del Sur, Cuba. Poeyana 288: 1-17.

Perera, A. 1985. Datos sobre la dieta de Cyclura nubila (Sauria: Iguanidae) en los alrededores de Cayo Largo del Sur, Cuba. Poeyana 291: 1-12.

Perez-Buitrago, N.F.; Alvarez, A.O.; Garcia, M.A. 2006. Cannibalisim in an introduced population of Cyclura nubila nubila on Isla Magueyes, Puerto Rico. Iguana (West Hartford) 13(3): 206-208.

Petzold, H.G. 1962. Aus Zoologischen Gärten. Über den Kuba-Leguan. Aquarien Terrarien 9: 128-129.

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Schmidt, J. 1997. Erfahrungen bei der Zucht von Cyclura-Arten. Iguana Rundschreiben 10(1)(17): 21-26.

Shaw, C.E. 1954. Captive-bred Cuban iguanas Cyclura macleayi macleayi. Herpetologica 10: 73-78.

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Cyclura onchiopsis

Powell, R. 1999. Herpetology of Navassa Island, West Indies. Caribbean Journal of Science 35(1-2): 1-13.

Powell, R. 2000. Cyclura onchiopsis. Catalogue of American Amphibians and Reptiles 710: 1-3.

Cyclura pinguis

Alberts, A.C. 2007. Behavioral considerations of headstarting as a conservation strategy for endangered Caribbean rock iguanas. Applied Animal Behaviour Science 102(3-4): 380-391.

Bradley, K.A.; Gerber, G. 2006. Release of headstarted iguanas in Anegada, British Virgin Islands. Re-introduction News 25: 14-16.

Carey, W.M. 1975. The rock iguana, Cyclura pinguis on Anegada, British Virgin Islands, with notes on Cyclura ricardi and C. cornuta. Bulletin of the Florida State Museum Biological Sciences 19(4): 189-233.

Cheek, C.A.; Hlavaty, S.; Mougey, K.; Perkins, R.N.; Peyton, M.A.; Ryan, C.N.; Zavaleta, J.C.; Boal, C.W.; Perry, G. 2013. Vertical structure use by the Stout Iguana (Cyclura pinguis) on Guana Island, BVI. IRCF Reptiles & Amphibians 20(3): 112–118.

Gerber, G.P. 2004. An update on the ecology and conservation of Cyclura pinguis on Anegada. Iguana (West Hartford) 11(1): 23-26.

Gerber, G.P.; Bradley, K. 2004. Anegada Iguana, Cyclura pinguis: 2003 update on field research and release project. Iguana (West Hartford) 11(1): 33.

Goodyear, N.C.; Lazell, J. 1994. Status of a relocated population of endangered Iguana pinguis on Guana Island, British Virgin Islands. Restoration Ecology 2(1): 43-50.

Lemm, J.M.; Steward, S.W.; Schmidt, T.F. 2005. Reproduction of the critically endangered Anegada Island iguana Cyclura pinguis at San Diego zoo. International Zoo Yearbook 39: 141-152.

LeVering, K.; Perry, G. 2003. Natural history notes: Cyclura pinguis (Stout Iguana, Anegada Rock Iguana). Juvenile predation. Herpetological Review 34(4): 367-368.

Mitchell, A.A.; Lau, J.; Chemnick, L.G.; Thompson, E.A.; Alberts, A.C.; Ryder, O.A.; Gerber, G.P. 2011. Using microsatellite diversity in wild Anegada iguanas (Cyclura pinguis) to establish relatedness in a captive breeding group of this critically endangered species. Conservation Genetics 12(3): 771-781.

Mitchell, N.C. 1999. Effect of introduced ungulates on density, dietary preferences, home range, and physical condition of the iguana (Cyclura pinguis) on Anegada. Herpetologica 55(1): 7-17.

Pagni, L.E. 2004. Public attitudes and perceptions regarding conservation of the Anegada Iguana, Cyclura pinguis: the use and role of social surveys in conservation. Iguana (West Hartford) 11(1): 34.

Perry, G.; Lazell, J.; LeVering, K.; Mitchell, N.C. 2007. Body size and timing of reproduction in the highly endangered stout iguana, Cyclura pinguis, in the British Virgin Islands. Caribbean Journal of Science 43(1): 155-159.

Perry, G.; LeVering, K.; Mitchell, N.C. 2003. Natural history notes: Cyclura pinguis (Stout Iguana, Anegada Rock Iguana). Juvenile behavior. Herpetological Review 34(4): 367.

Skipper, B.; Grisham, B.; Kalyvaki, M.; McGaughey, K.; Mougey, K.; Navarrete, L.; Rondeau, R.; Boal, C.; Perry, G. 2013. Non-overlapping distributions of Feral Sheep (Ovis aries) and Stout Iguanas (Cyclura pinguis) on Guana Island, British Virgin Islands. IRCF Reptiles & Amphibians 20(1): 7–15.

Werning, H. 2011. Einer der seltensten Leguane der Welt - der Anegada-Wirtelschwanzleguan, Cyclura pinguis. Iguana Rundschreiben 23(2) 2010[2011]: 13-25.

Cyclura ricordii

Arias, Y.; Incháustequi, S.; Rupp, E. 2004. Cyclura ricordii on the Barahona Peninsula: a preliminary report. Iguana (West Hartford) 11(1): 8-14.

Banbury, B.L.; Ramos, Y.M. 2005. The rock iguanas of Parque Nacional Isla Cabritos. Iguana (West Hartford) 12(4): 256-261.

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Harmon, K. 1998. Plants, iguanas and islands: a zoo horticulturist visits Hispaniola. pp. 249-251. In: Anonymous. Proceedings of the 1998 joint conference AAZK EMA AZH, Indianapolis 1998. Indianapolis Zoological Society, Inc., Indianapolis. 286 pp.

Hartley, L.M.; Glor, R.E.; Sproston, A.L.; Powell, R.; Parmerlee, J.S. 2000. Germination rates of seeds consumed by two species of rock iguanas (Cyclura spp.) in the Dominican Republic. Caribbean Journal of Science 36(1-2): 149-151.

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Ramer, J. 2004. A survey of Ricord's Iguanas (Cyclura ricordii) and Rhinoceros Iguanas (Cyclura cornuta cornuta) in Isla Cabritos National Park, Dominican Republic 2003: a preliminary report. Iguana (West Hartford) 11(2): 88-97.

Ramer, J. 2004. Ricord's Iguana, Cyclura ricordii: Isla Cabritos population census 2003. Iguana (West Hartford) 11(1): 34-35.

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Cyclura rileyi

Carter, R.L.; Hayes, W.K. 2004. Conservation of an endangered Bahamian rock iguana, II: morphological variation and conservation priorities. pp. 258-273. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

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Cyclura stejnegeri

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Pérez, N.; McMillan, O.; Alvarez, A.O.; García, M.A. 2004. Territorial and reproductive behavior of the Mona Rock Iguana, Cyclura cornuta stejnegeri. Iguana (West Hartford) 11(1): 32.

Perez-Buitrago, N.F.; Garcia, M.A.; Sabat, A.M.; Delgado, J.; Alvarez, A.; McMillan, O.; Funk, S.M. 2008. Do headstart programs work? Survival and body condition in headstarted Mona Island iguanas Cyclura cornuta stejnegeri. Endangered Species Research 6(1): 55-65.

Perez-Buitrago, N.F.; Sabat, A.M. 2007. Natal dispersal, home range and habitat use of hatchlings of the Mona Island iguana (Cyclura cornuta stejnegeri). Applied Herpetology 4(4): 365-376.

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Pérez-Buitrago, N.F.; Sabat, A.M.; Owen, M.W. 2010. Spatial ecology of the endangered Mona Island iguana Cyclura cornuta stejnegeri: does territorial behavior regulate density? Herpetological Monographs 24: 86-110.

Perotto-Baldivieso, H.L.; Melendez-Ackerman, E.; Garcia, M.A.; Leimgruber, P.; Cooper, S.M.; Martinez, A.; Calle, P.; Ramos Gonzales, O.M.; Quinones, M.; Christen, C.A.; Pons, G. 2009. Spatial distribution, connectivity, and the influence of scale: habitat availability for the endangered Mona Island rock iguana. Biodiversity and Conservation 18(4): 905-917.

Powell, R. 1999. Herpetology of Navassa Island, West Indies. Caribbean Journal of Science 35(1-2): 1-13.

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