Reptiles of the Calamian Islands [Philippines] Related checklists:  Philippines - Busuanga - Calauit - Coron - Culion - Sangat Bibliography of the reptiles of Calamian Islands Taxonomic history of the reptiles of Calamian Islands Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: April 14th, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus mindorensis Philippine Freshwater Crocodile Alcala 1986 Trutnau & Sommerlad 2006

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Gaulke 1996 Gaulke 1999 Hallermann 2005 Taylor 1922 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus semperi White-spotted Anglehead Agama Alcala 1986  Family Gekkonidae (Typical Geckos) Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Gaulke 1999 Genus Gekko (Tokays)   Gekko gecko Tokay Brown & Alcala 1978 Gaulke 1999   Gekko monarchus Spotted House Gecko Gaulke 1999 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Brown & Alcala 1978 Gaulke 1999   Hemidactylus platyurus Flat-tailed Gecko Gaulke 1999  Family Scincidae (Skinks) Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown & Alcala 1980 Brown 1991 Gaulke 1999 Genus Eutropis (South Asian Sun Skinks)   Eutropis indeprensa Mindoro Sun Skink Gaulke 1999   Eutropis multifasciata Many-lined Sun Skink Brown & Alcala 1980 Gaulke 1999 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink David, Pauwels, Lays & Lenglet 2006  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus palawanensis Palawan Monitor Gaulke 1996 [as marmoratus] Gaulke 1999 [as marmoratus] Koch, Auliya, Schmitz, Kuch & Boehme 2007 [as marmoratus] Koch, Gaulke & Boehme 2010 Lisle 1996

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Gaulke 1999 Leviton, Siler, Weinell & Brown 2018  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake David, Pauwels, Lays & Lenglet 2006 Gaulke 1999 Iskandar & Colijn 2001 Leviton 1967 Leviton, Siler, Weinell & Brown 2018 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga cynodon Dog-toothed Cat Snake Alcala 1986 Gaulke 1996 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus philippinus Palawan Rat Snake Alcala 1986 David, Pauwels, Lays & Lenglet 2006 [as erythrura] Gaulke 1996 [as erythrura] Gaulke 1999 Iskandar & Colijn 2001 Leviton 1979 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis caudolineatus Malayan Stripe-tailed Bronzeback Gaulke 1999 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018   Dendrelaphis marenae Philippine Painted Bronzeback Gaulke 1999 [as pictus] Iskandar & Colijn 2001 [as pictus] Leviton 1968 [as pictus] Leviton, Siler, Weinell & Brown 2018 Vogel & Rooijen 2008 Genus Dryophiops (Malayan Vine Snakes)   Dryophiops rubescens Keel-bellied Vine Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 Genus Liopeltis (Ringneck Snakes)   Liopeltis philippina Culion Ringneck Snake Alcala 1986 Gaulke 1999 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Som, Grismer, Wood, Quah, Brown, Diesmos, Weinell & Stuart 2020 Genus Oligodon (Kukri Snakes)   Oligodon notospilus Palawan Kukri Snake Alcala 1986 [as vertebralis] Gaulke 1999 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018   * Oligodon perkinsi Culion Kukri Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Supsup & Carestia 2020 Genus Pseudorabdion (Dwarf Reed Snakes)   Pseudorabdion oxycephalum Sharp-nosed Dwarf Reed Snake Iskandar & Colijn 2001 Leviton & Brown 1959 Leviton 1963 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis chrysargos Speckled-bellied Keelback Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haas 1950 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis bivittatus Palawan Black-headed Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Calliophis (South Asian Coral Snakes)   Calliophis bilineatus Two-striped Coral Snake Alcala 1986 Brown, Smart, Leviton & Smith 2018 Gaulke 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Vogel 2006 Genus Naja (Typical Cobras)   Naja sumatrana Sunda Spitting Cobra Gaulke 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Sy, Dycoco, Rodriguez, Du, Wisco, Pangilinan & Banaguas 2016 Wuster & Thorpe 1990 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra David, Pauwels, Lays & Lenglet 2006 Gaulke, Demegillo, Reiter & Tacud 2003  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Heatwole, Busack & Cogger 2005  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python David, Pauwels, Lays & Lenglet 2006 Gaulke 1999 Leviton, Siler, Weinell & Brown 2018

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Gaulke 1987  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle David, Pauwels, Lays & Lenglet 2006 Diesmos, Brown, Alcala & Sison 2008 Genus Cyclemys (Asian Leaf Turtles)   Cyclemys dentata Common Leaf Turtle Diesmos, Brown, Alcala & Sison 2008 Fritz, Gaulke & Lehr 1997 Gaulke 1999