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Bibliography of the reptiles of
Pennsylvania  [USA]

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Allen, W.B. 1992. The snakes of Pennsylvania. Reptile and Amphibian Magazine, Pottsville, Pennsylvania. 33 pp.

Assetto, R. 1982. A list of reptiles and amphibians species collected randomly in lower Bucks County. Bulletin of the Philadelphia Herpetological Society 30: 15-18.

Atkinson, D.A. 1901. The reptiles of Alleghany County, Pennsylvania. Annals of the Carnegie Museum 1: 145-157.

Baldauf, R.J. 1943. Handlist of Berks County amphibians and reptiles. Leaflets Mengel Nat. Hist. Soc. Penn. 1: 1-8.

Bauer, A.M.; Witwer, M. 1995. The herpetofauna of Nottingham Park and the Serpentine Barrens of Chester County, Pennsylvania. Bulletin of the Chicago Herpetological Society 30(1): 3-9.

Bell, E.L. 1957. Some herpetological records from Central Pennsylvania. Herpetologica 13: 138-139.

Bell, E.L. 1960. Observations on the amphibians and reptiles of Huntingdon County, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 34: 159-161.

Brett, J.J. 1991. The Mountain and the migration: a guide to Hawk Mountain. Cornell University Press, Ithaca. 114 pp.

Brittingham, M.C.; Goodrich, L.J. 2010. Wildlife habitat: the key to abundance, distribution, and diversity. pp. 10-19. In: Steele, M.A.; Brittingham, M.C.; Maret, T.J.; Merritt, J.F. (eds.). Terrestrial vertebrates of Pennsylvania: a complete guide to species of conservation concern. Johns Hopkins University Press, Baltimore. 507 pp.

Chambers, D.L. 2006. An updated list of the amphibians and reptiles of Indiana County, Pennsylvania (USA). Journal of the Pennsylvania Academy of Science 80(1): 12-17.

Clements, D. 1986. June PHS field survey. Philadelphia Herpetological Society Newsletter 1986: 2.

Conant, R. 1942. Amphibians and reptiles from Dutch Mountain (Pennsylvania) and vicinity. American Midland Naturalist 27(1): 154-170.

Conant, R. 1959. Lacerta colony still extant at Philadelphia. Copeia 1959: 335-336.

Cypher, E.A.; Yahner, R.H.; Storm, G.L.; Cypher, B.L. 1986. Flora and fauna survey in a proposed recreational area of Valley Forge National Historical Park. Proceedings of the Pennsylvania Academy of Science 60(1): 47-50.

Delis, P.R.; Kindlin, C.; Stewart, R.L. 2010. The herpetofauna of Letterkenny army depot, south-central Pennsylvania: a starting point to the long-term monitoring and management of amphibians and reptiles. Journal of Kansas Herpetology 34: 11-16.

Derge, K.L.; Yahner, R.H.; Mravintz, J. 2001. Survey of reptiles and amphibians in Gettysburg National Park with reference to Michaux State Forest. Journal of the Pennsylvania Academy of Science 75(2-3): 43-49.

Ernst, C.H. 1967. Notes on the herpetofauna of the White Oak Bird Sanctuary. Bulletin of the Philadelphia Herpetological Society 15: 32-36.

Feuer, J.; Feuer, R.C. 1977. Imperilled amphibians and reptiles of Pennsylvania and New Jersey. Bulletin of the Philadelphia Herpetological Society 25: 26-28.

Feuer, R.C. 1967. Key to the adult amphibians and reptiles of Pennsylvania. Bulletin of the Philadelphia Herpetological Society 15: 7-17.

Gray, B.S. 2002. Observations on the herpetofauna of Asbury Woods Greenway in Pennsylvania. Bulletin of the Chicago Herpetological Society 37(2): 21-24.

Gray, B.S. 2006. The amphibians and reptiles of the Asbury Woods Greenway, Erie County, Pennsylvania. Bulletin of the Maryland Herpetological Society 42(2): 115-126.

Gray, B.S. 2009. Recent observations of the herpetofauna of a former national superfund site in Erie, Pennsylvania. Journal of Kansas Herpetology 31: 9-11.

Gray, B.S.; Lethaby, M. 2008. The amphibians and reptiles of Erie County, Pennsylvania. Bulletin of the Maryland Herpetological Society 44(2): 49-69.

Groves, J.D. 1977. Priorities for effective management of the reptile fauna in Pennsylvania. Bulletin of the Philadelphia Herpetological Society 25: 15-17.

Harding, J. 2000. Reptiles and Amphibians of the Great Lakes Region. University of Michigan Press, Ann Arbor. 378 pp.

Hartzell, S.M. 2017. New township records for amphibians and reptiles in Columbia County, Pennsylvania, USA. Bulletin of the Chicago Herpetological Society 52(10): 173-175.

Heilman, R.A. 1951. A list of the amphibians and reptiles of Lebanon County, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 25: 44-46.

Hudson, R.G. 1954. An annotated list of the reptiles and amphibians of the Unami Valley, Pennsylvania. Herpetologica 10: 07-72.

Hughes, D.F.; Delis, P.R.; Meshaka, W.E. 2018. Studies of a snake assemblage from artificial ponds at an activemilitary site in Pennsylvania. Herpetology Notes 11: 35-48.

Hulse, A.C.; Hulse, K.L. 1992. New county records for amphibians and reptiles from Pennsylvania. Herpetological Review 23(2): 62-64.

Hulse, A.C.; McCoy, C.J.; Censky, E.J. 2001. Amphibians and reptiles of Pennsylvania and the northeast. Cornell University Press, Ithaca, New York. 419 pp.

Knepp, T.H. 1941. Some additions to the list of amphibians and reptiles of Bedford County. Proceedings of the Pennsylvania Academy of Science 15: 164.

Lovich, J.E. 1997. Additional records of amphibians and reptiles from Indiana County, Pennsylvania. Journal of the Pennsylvania Academy of Science 71(1): 35-38.

Lovich, J.E.; Ernst, C.H.; Ernst, E.M.; Riley, J.L. 2014. A 21-year study of seasonal and interspecific variation of hatchling emergence in a Nearctic freshwater turtle community: to overwinter or not to overwinter? Herpetological Monographs 28: 93-109.

Maret, T.J. 2010. The amphibians and reptiles. pp. 26-105. In: Steele, M.A.; Brittingham, M.C.; Maret, T.J.; Merritt, J.F. (eds.). Terrestrial vertebrates of Pennsylvania: a complete guide to species of conservation concern. Johns Hopkins University Press, Baltimore. 507 pp.

McCoy, C.J. (ed.) 1985. Species of special concern in Pennsylvania. Chapter 4 - amphibians and reptiles. Carnegie Museum of Natural History Special Publication 11: 258-295.

McCoy, C.J. 1982. Amphibians and reptiles in Pennsylvania. Carnegie Museum of Natural History Special Publication 6: 1-91.

McCoy, C.J. 1986. Bibliography of Pennsylvanian herpetology: 1981-1986. Proceedings of the Pennsylvania Academy of Science 60(2): 122-124.

McCoy, C.J. 1986. New perspectives on threatened and endangered amphibians and reptiles in Pennsylvania. pp. 202-210. In: Majumdar, S.K.; Brenner, F.J. & Rhoads, A.F. (eds.). Endangered and threatened species programs in Pennsylvania and other states: causes, issues and management. Pennsylvania Academy of Science, Pennsylvania, USA. 519 pp.

McCoy, C.J. 1989. The amphibians and reptiles of Pennsylvania wetlands: an ecological survey. pp. 171-182. In: Majumdar, S.K.; Brooks, R.P.; Brenner, F.J. & Tiner, R.W., Jr (eds.). Wetlands ecology and conservation: emphasis in Pennsylvania. Pennsylvania Academy of Science Publications, Pennsylvania. 394 pp.

McKinstry, D.M. 1975. Notes on the herpetology of Presque Isle State Park, Erie, Pennsylvania. Bulletin of the Maryland Herpetological Society 11(1): 21-26.

McKinstry, D.M.; Felege, S. 1974. Snakes of northwestern Pennsylvania. Bulletin of the Maryland Herpetological Society 10(1): 29-31.

McKinstry, D.M.; Lethaby, M.A.; Cunningham, H.N. 1991. Amphibians and reptiles of Presque Isle State Park, Erie County, Pennsylvania. Journal of the Pennsylvania Academy of Science 65(1): 17-23.

Meshaka, W.E. 2010. Field observations of eleven species of snakes during twenty-six years at Hawk Mountain Sanctuary, Pennsylvania. Journal of Kansas Herpetology 36: 17-20.

Meshaka, W.E. 2010. Seasonal activity and breeding seasons of snakes from Powdermill Nature Reserve in western Pennsylvania: the importance of site-specific data in land management programs. Herpetological Conservation and Biology 5(1): 155-165.

Meshaka, W.E.; Marshall, S.D.; Guiher, T.J.; Zemba, L. 2009. Grassland snake assemblages in central and western Pennsylvania and northeastern Ohio, USA. Herpetological Bulletin 110: 8-19.

Meshaka, W.E.; Netting Huff, J.; Leberman, R.C. 2008. Amphibians and reptiles of Powdermill Nature Reserve in western Pennsylvania. Journal of Kansas Herpetology 25: 12-18.

Netting, M.G. 1930. The occurrence of lizards in Pennsylvania. Annals of the Carnegie Museum 19(3): 169-174.

Netting, M.G. 1930. The poisonous snakes of Pennsylvania and the treatment of snakebites. Annals of the Carnegie Museum 19(3): 175-184.

Netting, M.G. 1936. Hand list of the amphibians and reptiles of Pennsylvania. Carnegie Mus. Pennsylvania Herpetology Leaflet 1: Unpaginated.

Netting, M.G. 1936. The amphibians and reptiles of Indiana County, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 10: 25-28.

Netting, M.G. 1940. Handlist of the amphibians and reptiles of Pennsylvania. Bienn. Rept. Pennsylvan. Fish. Comm. for 1936-1938 1939: 109-134.

Netting, M.G. 1946. The amphibians and reptiles of Pennsylvania. Board Fish. Comm., Harrisburg, Pa. 1946: 1-29.

Pawling, R.O. 1939. The amphibians and reptiles of Union County, Pennsylvania. Herpetologica 1939(1): 165-169.

Reinert, H.K. 1990. A profile and impact assessment of organized rattlesnake hunts in Pennsylvania. Journal of the Pennsylvania Academy of Science 64(3): 136-144.

Richmond, N.D. 1955. Poisonous snakes of Pennsylvania, how to recognise them. Carnegie Mus. Conservation Folder. 4 pp.

Ross, B.; Yahner, R.H.; Karish, J.F.; Kubel, J.E. 2007. Survey of amphibians and reptiles at Allegheny Portage Railroad National Historic Site and Johnstown Flood National Memorial, Pennsylvania. Journal of the Pennsylvania Academy of Science 81(1): 3-7.

Senecal Kearney, C. (ed.) 1981. Pennsylvania species of special concern. Endangered Species Technical Bulletin 6(11): 4-7.

Smith, A.G. 1945. The amphibians and reptiles of Westmoreland County, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 19: 72-80.

Steele, M.A.; Brittingham, M.C.; Maret, T.J.; Merritt, J.F. (eds.). 2010. Terrestrial vertebrates of Pennsylvania: a complete guide to species of conservation concern. Johns Hopkins University Press, Baltimore. 507 pp.

Steele, M.A.; Reeder, D.M.; Maret, T.J.; Brittingham, M.C. 2010. Critical and emerging issues in the conservation of terrestrial vertebrates. pp. 390-398. In: Steele, M.A.; Brittingham, M.C.; Maret, T.J.; Merritt, J.F. (eds.). Terrestrial vertebrates of Pennsylvania: a complete guide to species of conservation concern. Johns Hopkins University Press, Baltimore. 507 pp.

Steiner, R.P. 1980. Some effects of a controlled spring burn on vertebrate populations of the Blazing Star Prairie, Jennings Nature Reserve, Butler Co., Pa. (Abstract). Proceedings of the Pennsylvania Academy of Science 54(1): 103.

Stewart, G.D.; Grant, R.R. 1963. County records for Pennsylvania and New Jersey. Bulletin of the Philadelphia Herpetological Society 11(3-4): 57.

Stewart, N.H. 1928. Some rare vertebrates of the Susquehanna Valley, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 2: 21-24.

Stone, W. 1906. Notes on reptiles and batrachians of Pennsylvania, New Jersey, and Delaware. American Naturalist 40: 159-170.

Stratford, J.; Steele, M.A. 2010. Conservation of terrestrial vertebrates: an overview and synthesis. pp. 1-9. In: Steele, M.A.; Brittingham, M.C.; Maret, T.J.; Merritt, J.F. (eds.). Terrestrial vertebrates of Pennsylvania: a complete guide to species of conservation concern. Johns Hopkins University Press, Baltimore. 507 pp.

Surface, H.A. 1906. The serpents of Pennsylvania. Pennsylvania Dept. Agric. Zool. Bull. Div. Zool. 4: Unpaginated.

Surface, H.A. 1908. First report on the economy of Pennsylvania turtles. Pennsylvania Dept. Agric. Zool. Bull. Div. Zool. 6: 107-195.

Surface, H.A. 1908. First report on the lizards of Pennsylvania. Pennsylvania Dept. Agric. Zool. Bull. Div. Zool. 5 [1907]: 236-258.

Swanson, P.L. 1948. Notes on the amphibians of Venango County, Pennsylvania. American Midland Naturalist 40(2): 362-371.

Swanson, P.L. 1952. The reptiles of Venango County, Pennsylvania. American Midland Naturalist 47: 161-182.

Warny, P.R. 1985. A preliminary herpetological survey in the Serpentine Barrens of Chester County, Pennsylvania. Bulletin of the Chicago Herpetological Society 20(1): 23-24.

Williams, L. 2010. Pennsylvania's Wildlife Action Plan. pp. 20-25. In: Steele, M.A.; Brittingham, M.C.; Maret, T.J.; Merritt, J.F. (eds.). Terrestrial vertebrates of Pennsylvania: a complete guide to species of conservation concern. Johns Hopkins University Press, Baltimore. 507 pp.

Yahner, R.H.; Piergallini, N.H.; Ross, B.D. 2001. Survey of amphibians and reptiles in two types of managed forests in central Pennsylvania. Journal of the Pennsylvania Academy of Science 74(2-3): 48-51.