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Taxonomy of the genus Phymaturus Biodiversity of the family Liolaemidae

Bibliography of the genus
Phymaturus (South American Chuckwallas)

(Reptilia: Sauria: Liolaemidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Phymaturus in general

Andrade-Díaz, M.S.; Hibbard, T.N.; Díaz-Gómez, J.M. 2017. Identifying endemism areas: an example using Neotropical lizards. South American Journal of Herpetology 12(1): 61–75

Ávila, L.J.; Pérez, C.H.F.; Pérez, D.R.; Morando, M. 2011. Two new mountain lizard species of the Phymaturus genus (Squamata: Iguania) from northwestern Patagonia, Argentina. Zootaxa 2924: 1-21.

Boretto, J.M.; Ibargüengoytía, N.R. 2009. Phymaturus of Patagonia, Argentina: reproductive biology of Phymaturus zapalensis (Liolaemidae) and a comparison of sexual dimorphism within the genus. Journal of Herpetology 43(1): 96-104.

Cei, J.M.; Castro, L.P. 1973. Taxonomic and serological researches on the Phymaturus patagonicus complex. Journal of Herpetology 7(3): 237-247.

Cei, J.M.; Lescure, J. 1985. Identite de Lacerta palluma Molina, 1782, et revalidation de Centrura flagellifer Bell, 1843 (Reptilia, Sauria). Bulletin du Museum National d'Histoire Naturelle Section A Zoologie Biologie et Ecologie Animales 7(2): 451-459.

Cei, J.M.; Scolaro, J.A. 2006. The neotype of the type species of the Neotropical iguanian genus Phymaturus: a critical commentary on a recent opinion of the International Commission on Zoological Nomenclature. Zootaxa 1297: 17-22.

Corbalán, V.; Debandi, G.; Scolaro, J.A.; Ojeda, A.A. 2016. DNA barcoding of Phymaturus lizards reveals conflicts in species delimitation within the patagonicus clade. Journal of Herpetology 50(4): 654–666.

Cruz, F.B.; Belver, L.C.; Acosta, J.C.; Villavicencio, H.J.; Blanco, G.; Canovas, M.G. 2009. Thermal biology of Phymaturus lizards: evolutionary constraints or lack of environmental variation? Zoology (Jena) 112(6): 425-432.

Debandi, G.; Corbalan, V.; Scolaro, J.A.; Roig-Junent, S.A. 2012. Predicting the environmental niche of the genus Phymaturus: are palluma and patagonicus groups ecologically differentiated. Austral Ecology 37(3): 392-400.

Diaz Gomez, J.M. 2009. Historical biogeography of Phymaturus (Iguania: Liolaemidae) from Andean and patagonian South America. Zoologica Scripta 38(1): 1-7.

Espinoza, R.E. 2003. Comment on the proposed conservation of usage of the names Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma; Reptilia, Sauria) by designation of a neotype for Lacerta palluma Molina, 1782. Bulletin of Zoological Nomenclature 60(1): 58.

Etheridge, R.E. 1995. Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates 3142: 1-34.

Etheridge, R.E.; Savage, J.M. 2003. Case 3225. Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma; Reptilia, Sauria): Proposed conservation of usage of the names by designation of a neotype for Lacerta palluma Molina, 1782. Bulletin of Zoological Nomenclature 60(1): 38-41.

Etheridge, R.E.; Savage, J.M. 2004. Comments on the proposed conservation of usage of the names Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma Reptilia, Sauria) by designation of a neotype for Lacerta palluma. Case 3225. Bulletin of Zoological Nomenclature 61(1): 47.

Frost, D. 2004. Comments on the proposed conservation of usage of the names Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma Reptilia, Sauria) by designation of a neotype for Lacerta palluma. Case 3225. Bulletin of Zoological Nomenclature 61(1): 47.

González-Gutiérrez, P.A. 2016. Registro de una población de Phymaturus (Squamata, Liolaemidae) en los alrededores de Farellones, Región Metropolitana. Boletín Chileno de Herpetología 3: 28-29.

Grosso, J.R.; Cardozo, D.; Baldo, D.; Lobo, F. 2017. Multiple sex chromosome system and Robertsonian rearrangements involved in the chromosome evolution of the Phymaturus palluma group (Iguania: Liolaemidae). Journal of Herpetology 51(1): 154–160.

Halloy, M.; Robles, C.; Salica, M.J.; Semhan, R.V.; Juarez Heredia, V.I.; Natalin, V. 2013. Aportes en el estudio del comportamiento y ecología de lagartijas de los géneros Liolaemus y Phymaturus (Iguania: Liolaemini). Cuadernos de Herpetologia 27(1-2): 15-26.

Ibargüengoytía, N.R.; Acosta, J.C.; Boretto, J.M.; Villavicencio, H.J.; Marinero, J.A.; Krenz, J.D. 2008. Field thermal biology in Phymaturus lizards: comparisons from the Andes to the Patagonian steppe in Argentina. Journal of Arid Environments 72(9): 1620-1630.

International Commission on Zoological Nomenclature. 2005. Opinion 2118 (Case 3225). Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma, Reptilia, Sauria): usage of the names conserved by the designation of a neotype for Lacerta palluma Molina, 1782. Bulletin of Zoological Nomenclature 62(2): 116-117.

Lescure, J.; Cei, J.M. 1991. L'espece-type du genre Phymaturus Gravenhorst, 1838 (Reptilia, Sauria). Museo Regionale di Scienze Naturali Bollettino (Turin) 9(1): 173-175.

Lobo, F.; Abdala, C.S.; Valdecantos, S. 2012. Morphological diversity and phylogenetic relationships within a South-American clade of iguanian lizards (Liolaemidae: Phymaturus). Zootaxa 3315: 1-41.

Lobo, F.; Barrasso, D.A.; Hibbard, T.; Basso, N.G. 2016. On the evolution and diversification of an Andean clade of reptiles: combining morphology and DNA sequences of the palluma group (Liolaemidae: Phymaturus). Zoological Journal of the Linnean Society 176(3): 648-673.

Lobo, F.; Barrasso, D.A.; Pa, M.; Basso, N.G. 2018. Phylogenetic relationships within a patagonian clade of reptiles (Liolaemidae: Phymaturus) based on DNA sequences and morphology. Journal of Zoological Systematics and Evolutionary Research 56(4): 549-569.

Lobo, F.; Etheridge, R.E. 2013. The type locality of Phymaturus palluma (Molina, 1782) (Reptilia: Iguania: Liolaemidae) and the status of Phymaturus adrianae Pereyra, 1992 and other unnamed populations. South American Journal of Herpetology 8(2): 89-101.

Lobo, F.; Quinteros, S. 2005. A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia (São Paulo) 45(13): 143-177.

Lobo, F.; Quinteros, S. 2005. Taxonomic studies of the genus Phymaturus (Iguania: Liolaemidae): Redescription of Phymaturus patagonicus Koslowsky 1898, and revalidation and redescription of Phymaturus spurcus Barbour 1921. Journal of Herpetology 39(4): 533-540.

Morando, M.; Ávila, L.J.; Pérez, C.H.F.; Hawkins, M.A.; Sites, J.W. 2013. A molecular phylogeny of the lizard genus Phymaturus (Squamata, Liolaemini): implications for species diversity and historical biogeography of southern South America. Molecular Phylogenetics and Evolution 66(3): 694-714.

Moro, S.; Abdala, V. 1998. Cranial myology of some species of Liolaemus and Phymaturus (Squamata: Tropiduridae: Liolaeminae). Amphibia-Reptilia 19(2): 171-192.

Nunez, H.; Veloso, A.; Espejo, P.; Veloso, C.; Cortes, A.; Araya, S. 2010. Nuevas especies de Phymaturus (grupo "palluma") para la Zona Cordillerana Central de Chile (Reptilia, Sauria, Liolaemidae). Boletin del Museo Nacional de Historia Natural Chile 59: 41-74.

Pincheira-Donoso, D.; Fox, S.F.; Scolaro, J.A.; Ibargüengoytía, N.R.; Acosta, J.C.; Corbalan, V.; Medina, M.; Boretto, J.; Villavicencio, H.J.; Hodgson, D.J. 2011. Body size dimensions in lizard ecological and evolutionary research: exploring the predictive power of mass estimation equations in two Liolaemidae radiations. Herpetological Journal 21(1): 35-42.

Scolaro, J.A.; Pincheira-Donoso, D. 2010. Lizards at the end of the world: two new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) revealed in southern Patagonia of Argentina. Zootaxa 2393: 17-32.

Smith, H.M. 2003. Comment on proposed conservation of the usage of the names Phymaturus Gravenhorst, 1838 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma; Reptilia, Sauria) by designation of a neotype for Lacerta palluma Molina, 1782. Bulletin of Zoological Nomenclature 60(3): 220.

Troncoso-Palacios, J.; Ferri-Yáñez, F.; Laspiur, A.; Aguilar, C. 2018. An updated phylogeny and morphological study of the Phymaturus vociferator clade (Iguania: Liolaemidae). Zootaxa 4441(3): 447–466.

Valdecantos, S.; Lobo, F. 2015. First report of hemiclitores in females of South American Liolaemid lizards. Journal of Herpetology 49(2): 291-294.

Veloso, A.; Nunez, H.; Cei, J.M. 2003. Comment on the proposed conservation of usage of the names Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma Reptilia, Sauria) by designation of a neotype for Lacerta palluma. Case 3225. Bulletin of Zoological Nomenclature 60(4): 313-314.

Phymaturus aguanegra

Lobo, F.; Laspiur, A.; Acosta, J.C. 2013. Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from northwestern Argentina. Zootaxa 3683(2): 117-132.

Phymaturus aguedae

Troncoso-Palacios, J.; Esquerré, D. 2014. A new species of Phymaturus of the P. mallimaccii group from the Andes of central Chile (Iguania: Liolaemidae). Phyllomedusa 13(1): 3-15.

Phymaturus alicahuense

Nunez, H.; Veloso, A.; Espejo, P.; Veloso, C.; Cortes, A.; Araya, S. 2010. Nuevas especies de Phymaturus (grupo "palluma") para la Zona Cordillerana Central de Chile (Reptilia, Sauria, Liolaemidae). Boletin del Museo Nacional de Historia Natural Chile 59: 41-74.

Phymaturus antofagastensis

Acosta, J.C.; Villavicencio, H.J.; Blanco, G. 2008. Natural history notes: Phymaturus antofagastensis (NCN). Diet. Herpetological Review 39(1): 91.

Boretto, J.M.; Ibargüengoytía, N.R. 2006. Asynchronous spermatogenesis and biennial female cycle of the viviparous lizard Phymaturus antofagastensis (Liolaemidae): reproductive responses to high altitudes and temperate climate of Catamarca, Argentina. Amphibia-Reptilia 27(1): 25-36.

Phymaturus bibronii

Nunez, H.; Veloso, A.; Espejo, P.; Veloso, C.; Cortes, A.; Araya, S. 2010. Nuevas especies de Phymaturus (grupo "palluma") para la Zona Cordillerana Central de Chile (Reptilia, Sauria, Liolaemidae). Boletin del Museo Nacional de Historia Natural Chile 59: 41-74.

Troncoso-Palacios, J.; Lobo, F.; Etheridge, R.E.; Acosta, J.C.; Laspiur, A. 2013. The rediscovery of Oplurus bibronii Guichenot, 1848 a valid species of the liolaemid genus Phymaturus (Iguania: Liolaemidae). Zootaxa 3652(5): 547-561.

Phymaturus calcogaster

González-Marín, A.; Morando, M.; Avila, L.J. 2016. Morfología lineal y geométrica en un grupo de lagartijas patagónicasdel género Phymaturus (Squamata: Liolaemini). Revista Mexicana de Biodiversidad 87(2): 399-408.

Scolaro, J.A.; Cei, J.M. 2003. Una excepcional nueva especie de Phymaturus de la precordillera de Chubut, Argentina (Liolaemidae, Iguania, Lacertilia, Reptilia). Facena 19: 157-163.

Scolaro, J.A.; Tapiri, F.O.; Gonzalez, C. 2005. Phymaturus calcogaster: rectificacion de la localidad tipo y descripcion de la hembra (Reptilia, Iguania, Liolaemidae). Facena 21: 29-36.

Phymaturus camilae

Scolaro, A.J.; Jara, M.; Pincheira-Donoso, D. 2013. The sexual signals of speciation? A new sexually dimorphic Phymaturus species of the patagonicus clade from Patagonia Argentina. Zootaxa 3722(3): 317-332.

Phymaturus castillensis

Scolaro, J.A.; Pincheira-Donoso, D. 2010. Lizards at the end of the world: two new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) revealed in southern Patagonia of Argentina. Zootaxa 2393: 17-32.

Phymaturus ceii

Scolaro, J.A.; Ibargüengoytía, N.R. 2007. A new species of Phymaturus from rocky outcrops in the central steppe of Rio Negro province, Patagonia Argentina (Reptilia: Iguania: Liolaemidae). Zootaxa 1524: 47-55.

Phymaturus curivilcun

Scolaro, J.A.; Corbalán, V.; Tappari, O.F.; Streitenberger, L.O. 2016. Lizards at the end of the world: a new melanic species of Phymaturus of the patagonicus clade from rocky outcrops in the northwestern steppe of Chubut province, Patagonia, Argentina (Reptilia: Iguania: Liolaemidae). Boletin del Museo Nacional de Historia Natural Chile 65: 137-152.

Phymaturus damasense

Troncoso-Palacios, J.; Lobo, F. 2012. A new species of Phymaturus (Iguania: Liolaemidae) of the palluma group from Central Chile. Cuadernos de Herpetologia 26(2): 69-78.

Phymaturus darwini

Nunez, H.; Veloso, A.; Espejo, P.; Veloso, C.; Cortes, A.; Araya, S. 2010. Nuevas especies de Phymaturus (grupo "palluma") para la Zona Cordillerana Central de Chile (Reptilia, Sauria, Liolaemidae). Boletin del Museo Nacional de Historia Natural Chile 59: 41-74.

Phymaturus delheyi

Ávila, L.J.; Pérez, C.H.F.; Pérez, D.R.; Morando, M. 2011. Two new mountain lizard species of the Phymaturus genus (Squamata: Iguania) from northwestern Patagonia, Argentina. Zootaxa 2924: 1-21.

Phymaturus denotatus

Lobo, F.; Nenda, S.J.; Slodki, D. 2012. A new lizard of Phymaturus (Iguania: Liolaemidae) from Argentina. Herpetologica 68(1): 121-133.

Phymaturus desuetus

Scolaro, J.A.; Tappari, O.F. 2009. Una nueva especie del género Phymaturus del “grupo patagonicus” en los afLoramientos rocosos del sudoeste de la provincia de Rio Negro, Patagonia Argentina (Reptilia: Iguania: Liolaemidae). Naturalia Patagónica 4: 81–94.

Phymaturus dorsimaculatus

Lobo, F.; Quinteros, S. 2005. A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia (São Paulo) 45(13): 143-177.

Phymaturus etheridgei

Lobo, F.; Abdala, C.S.; Valdecantos, S. 2010. Taxonomic studies of the genus phymaturus (Iguania: Liolaemidae): description of four new species. South American Journal of Herpetology 5(2): 102-126.

Phymaturus excelsus

Lobo, F.; Quinteros, S. 2005. A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia (São Paulo) 45(13): 143-177.

Phymaturus extrilidus

Cabezas-Cartes, F. 2013. Natural history notes: Phymaturus spectabilis. Predation. Herpetological Review 44(4): 679.

Lobo, F.; Espinoza, R.E.; Sanabria, E.A.; Quiroga, L.B. 2012. A new Phymaturus (Iguania: Liolaemidae) from the southern extreme of the Argentine Puna. Copeia 2012(1): 12-22.

Pérez, M.A.; Borghi, C.E.; Beninato, V.A. 2013. Natural history notes: Phymaturus extrilidus. Predation. Herpetological Review 44(4): 679.

Phymaturus felixi

Lobo, F.; Abdala, C.S.; Valdecantos, S. 2010. Taxonomic studies of the genus phymaturus (Iguania: Liolaemidae): description of four new species. South American Journal of Herpetology 5(2): 102-126.

Phymaturus gynechlomus

Corbalan, V.; Scolaro, J.A.; Debandi, G. 2009. A new species of the genus Phymaturus of the flagellifer group from central-western Mendoza, Argentina (Reptilia: Iguania: Liolaemidae). Zootaxa 2021: 42-56.

Scolaro, J.A. 2010. Redescripcion del neotipo de Phymaturus palluma: un aporte a la delimitacion de su tierra tipica (Reptilia, Sauria, Liolaemidae). Boletin del Museo Nacional de Historia Natural Chile 59: 29-39.

Phymaturus indistinctus

Minoli, I.; Morando, M.; Avila, L.J. 2015. Reptiles of Chubut province, Argentina: richness, diversity, conservation status and geographic distribution maps. ZooKeys 498: 103-126.

Phymaturus laurenti

Lobo, F.; Abdala, C.S.; Valdecantos, S. 2010. Taxonomic studies of the genus phymaturus (Iguania: Liolaemidae): description of four new species. South American Journal of Herpetology 5(2): 102-126.

Phymaturus loboi

Troncoso-Palacios, J.; Ferri-Yáñez, F.; Laspiur, A.; Aguilar, C. 2018. An updated phylogeny and morphological study of the Phymaturus vociferator clade (Iguania: Liolaemidae). Zootaxa 4441(3): 447–466.

Phymaturus mallimaccii

Cei, J.M. 1980. New endemic iguanid lizards from the Famatina Mountains of western Argentina. Journal of Herpetology 14(1): 57-64.

Phymaturus manuelae

Scolaro, J.A.; Ibargüengoytía, N.R. 2008. A new fragment for the understanding of the puzzling evolutive process of the Phymaturus genus: a new species of the patagonicus group from Patagonia, Argentina (Reptilia: Iguania: Liolaemidae). Zootaxa 1939: 38-50.

Phymaturus maulense

Nunez, H.; Veloso, A.; Espejo, P.; Veloso, C.; Cortes, A.; Araya, S. 2010. Nuevas especies de Phymaturus (grupo "palluma") para la Zona Cordillerana Central de Chile (Reptilia, Sauria, Liolaemidae). Boletin del Museo Nacional de Historia Natural Chile 59: 41-74.

Phymaturus nevadoi

Cei, J.M.; Roig, V.G. 1975. A new lizard from the Sierra del Nevado Mountains, central Argentina. Journal of Herpetology 9(2): 256.

Phymaturus palluma

Alzamora, A.; Gallardo, C.; Vukasovic, A.; Thomson-S., R.; Camousseigt, N.; Charrier, A.; Garin, C.; Lobos, G. 2010. Natural history notes: Phymaturus flagellifer (Matuasto). Brumation behavior. Herpetological Review 41(1): 85.

Barros, R.D.; Herrera, S.C. 1950. Reptiles de la Provincia de Santiago. Revista de la Academia Colombiana de Ciencias Exactas Fisicas y Naturales 7(28): 482-489.

Cartes, F.C.; Boretto, J.M.; Acosta, J.C.; Jahn, G.; Blanco, G.; Laspiur, A.; Ibargüengoytía, N.R. 2010. Reproductive biology of Phymaturus cf. palluma: a vulnerable lizard from the highlands of the Andes, San Juan, Argentina. Herpetological Conservation and Biology 5(3): 430-440.

Castro, S.A.; Laspiur, A.; Acosta, J.C. 2013. [Annual and intrapopulational variation in diet of Phymaturus cf. palluma (Iguania: Liolaemidae) from the Central Andes in Argentina]. (In Spanish). Revista Mexicana de Biodiversidad 84(4): 1258-1265.

Cei, J.M. 1971. Consideraciones sobre las relaciones taxinomicas de Phymaturus patagonicus y Phymaturus palluma. Acta Zoologica Lilloana 28: 37-46.

Cei, J.M. 1972. Consideraciones sobre las relaciones taxinómicas de Phymaturus patogonicus y Phymaturus palluma. Acta Zoologica Lilloana 28[1971]: 37-46.

Celedon-Neghme, C.; Salgado, C.R.; Victoriano, P.F. 2005. Preferencias alimentarias y potencial dispersor del lagarto herbivoro Phymaturus flagellifer (Tropiduridae) en los Andes. Gayana 69(2): 266-276.

Dacuik, J. 1979. Notas faunisticas y bioecologicas de Peninsula Valdes y Patagonia: 1. Datos bioecologicos obtenidos en cautividad del lagarto Phymaturus palluma palluma (Molina), 1782. (Lacert., Iguanidae). Anales de Parques Nacionales (Buenos Aires) 14: 80-86.

Eisenberg, T.; Werning, H. 2012. Phymaturus cf. palluma in captivity: observations on its reproduction and biology. Salamandra 48(4): 198-206.

Etheridge, R.E.; Savage, J.M. 2006. Stability restored to the name Phymaturus palluma: a response to Cei and Scolaro (2006). Herpetological Review 37(4): 406-407.

Habit, E.M.; Ortiz, J.C. 1994. Ambito de hogar de Phymaturus flagellifer (Reptilia, Tropiduridae). Boletin de la Sociedad de Biologia de Concepcion 65: 149-152.

Habit, E.M.; Ortiz, J.C. 1996. Ciclo reproductivo de Phymaturus flagellifer (Reptilia, Tropiduridae). Boletin de la Sociedad de Biologia de Concepcion 67: 7-14.

Habit, E.M.; Ortiz, J.C. 1996. Patrones de comportamiento y organizacion social de Phymaturus flagellifer (Reptilia, Tropiduridae). pp. 141-154. In: Pefaur, J.E. (ed.). Herpetologia Neotropical. Actas del II Congreso Latinoamericano de Herpetologia. II volumen. Universidad de los Andes, Consejo de Publicaciones, Merida. 451 pp.

Lamborot, M.; Navarro-Suarez, M. 1984. Karyotypes and sex determination in Phymaturus palluma Molina (Iguanidae). Herpetologica 40(3): 258-264.

Lobo, F.; Abdala, C.S.; Valdecantos, S. 2010. Taxonomic studies of the genus phymaturus (Iguania: Liolaemidae): description of four new species. South American Journal of Herpetology 5(2): 102-126.

Lobo, F.; Etheridge, R.E. 2013. The type locality of Phymaturus palluma (Molina, 1782) (Reptilia: Iguania: Liolaemidae) and the status of Phymaturus adrianae Pereyra, 1992 and other unnamed populations. South American Journal of Herpetology 8(2): 89-101.

Macola, G.S.; Castro, L.P.; Castro, M.P.; Mattiussi, M.B. 1985. Estudio comparativo del ciclo espermatogenico anual de las especies de una comunidad de saurios de altura, Uspallata-Las Heras-Mendoza (Sauria-Iguanidae). Revista del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" e Instituto Nacional de Investigacion de las Ciencias Naturales Zoologia 13: 139-145.

Pincheira-Donoso, D. 2002. Nota sobre una emision de sonidos en Phymaturus flagellifer (Bell, 1843) (Sauria - Tropiduridae - Liolaeminae). Noticiario Mensual Museo Nacional de Historia Natural (Santiago) 348: 8-9.

Schifter, H. 1965. Phymaturus palluma, ein chilenischer Hochgebirgsleguan. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 18: 342-344.

Scolaro, J.A. 2010. Redescripcion del neotipo de Phymaturus palluma: un aporte a la delimitacion de su tierra tipica (Reptilia, Sauria, Liolaemidae). Boletin del Museo Nacional de Historia Natural Chile 59: 29-39.

Smith, H.M. 2003. Comment on proposed conservation of the usage of the names Phymaturus Gravenhorst, 1838 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma; Reptilia, Sauria) by designation of a neotype for Lacerta palluma Molina, 1782. Bulletin of Zoological Nomenclature 60(3): 220.

Speroni, N.B. de; Cabrera, M.R.; Manca, L. 1984. Consideraciones sobre la osteologia craneal, hioides, esternon y lengua de Phymaturus palluma (Molina, 1782) (Sauria, Iguanidae). Historia Natural (Corrientes) 3(24): 197-212.

Vicenzi, N. 2015. Natural history notes: Phymaturus palluma (High Mountain Lizard). Coprophagy. Herpetological Review 46(1): 93.

Vicenzi, N.; Massarelli, R. 2018. Natural history notes: Phymaturus palluma (High Mountain Lizard). Diet. Herpetological Review 49(1): 121.

Vicenzi, N.; Massarelli, R.; Chamorro, E.; Barros, A. 2017. Highest record for the lizard Phymaturus palluma (Squamata: Liolaemidae) in Central Andes, Argentina. Herpetology Notes 10: 373-374.

Vidal, M.A.; Habit, E.; Victoriano, P.; González-Gajardo, A.; Ortiz, J.C. 2010. Thermoregulation and activity pattern of the high-mountain lizard Phymaturus palluma (Tropiduridae) in Chile. Zoologia 27(1): 13-18.

Phymaturus patagonicus

Avila, L.J.; Morando, M.; Perez Daniel, R. 1999. Geographic distribution: Phymaturus patagonica. Herpetological Review 30(3): 174.

Cei, J.M. 1971. Consideraciones sobre las relaciones taxinomicas de Phymaturus patagonicus y Phymaturus palluma. Acta Zoologica Lilloana 28: 37-46.

Cei, J.M. 1972. Consideraciones sobre las relaciones taxinómicas de Phymaturus patogonicus y Phymaturus palluma. Acta Zoologica Lilloana 28[1971]: 37-46.

Chiszar, D.; Gingery, T.; Gingery, B.; Smith, H.M. 1999. Natural history notes: Phymaturus patagonicus (Argentine Chuckwalla). Facultative parthenogenesis. Herpetological Review 30(2): 98.

Ibargüengoytía, N.R. 2004. Prolonged cycles as a common reproductive pattern in viviparous lizards from Patagonia, Argentina: reproductive cycle of Phymaturus patagonicus. Journal of Herpetology 38(1): 73-79.

Ibargüengoytía, N.R. 2005. Field, selected body temperature and thermal tolerance of the syntopic lizards Phymaturus patagonicus and Liolaemus elongatus (Iguania: Liolaemidae). Journal of Arid Environments 62(3): 435-448.

Lobo, F.; Quinteros, S. 2005. Taxonomic studies of the genus Phymaturus (Iguania: Liolaemidae): Redescription of Phymaturus patagonicus Koslowsky 1898, and revalidation and redescription of Phymaturus spurcus Barbour 1921. Journal of Herpetology 39(4): 533-540.

Piantoni, C.; Ibargüengoytía, N.R.; Cussac, V.E. 2006. Age and growth of the Patagonian lizard Phymaturus patagonicus. Amphibia-Reptilia 27(3): 385-392.

Pietrek, A.G.; Walker, R.S.; Novaro, A.J. 2009. Susceptibility of lizards to predation under two levels of vegetative cover. Journal of Arid Environments 73(4-5): 574-577.

Phymaturus payuniae

Corbalan, V.; Debandi, G. 2013. Basking behaviour in two sympatric herbivorous lizards (Liolaemidae: Phymaturus) from the Payunia volcanic region of Argentina. Journal of Natural History 47(19-20): 1365-1378.

Corbalan, V.; Debandi, G. 2014. Resource segregation in two herbivorous species of mountain lizards from Argentina. Herpetological Journal 24(4): 201-208.

Corbalan, V.; Debandi, G.; Kubisch, E. 2013. Thermal ecology of two sympatric saxicolous lizards of the genus Phymaturus from the Payunia region (Argentina). Journal of Thermal Biology 38(7): 384-389.

Phymaturus punae

Boretto, J.M.; Cabezas-Cartes, F.; Ibarguëngoytía, N.R. 2015. Energy allocation to growth and reproduction in a viviparous lizard endemic to the highlands of the Andes, Argentina. Journal of Zoology (London) 297(1): 77-86.

Boretto, J.M.; Fornes, M.W.; Jahn, G.A.; Acosta, J.C.; Ibargüengoytía, N.R. 2014. Testosterone cycle and regulation of reproductive events in the lizard Phymaturus punae (Liolaemidae) from the highlands of the Andes, Argentina. Journal of Herpetology 48(2): 172-180.

Boretto, J.M.; Ibargüengoytía, N.R.; Acosta, J.C.; Blanco, G.M.; Villavicencio, J.; Marinero, J.A. 2007. Reproductive biology and sexual dimorphism of a high-altitude population of the viviparous lizard Phymaturus punae from the Andes in Argentina. Amphibia-Reptilia 28(3): 427-432.

Cooper, W.E.; Habegger, J.J.; Espinoza, R.E. 2001. Responses to prey and plant chemicals by three iguanian lizards: Relationship to plants in the diet. Amphibia-Reptilia 22(3): 349-361.

Villavicencio, H.J.; Acosta, J.C.; Marinero, J.A. 2009. Natural history notes: Pristidactylus scapulatus (NCN). Saurophagy. Herpetological Review 40(2): 225-226.

Phymaturus querque

Lobo, F.; Abdala, C.S.; Valdecantos, S. 2010. Taxonomic studies of the genus phymaturus (Iguania: Liolaemidae): description of four new species. South American Journal of Herpetology 5(2): 102-126.

Phymaturus rahuensis

González-Marín, A.; Fulvio Pérez, C.H.; Minoli, I.; Morando, M.; Avila, L.J. 2016. A new lizard species of the Phymaturus patagonicus group (Squamata: Liolaemini) from northern Patagonia, Neuquén, Argentina. Zootaxa 4121(4): 412-430.

Phymaturus roigorum

Corbalan, V.; Debandi, G. 2013. Basking behaviour in two sympatric herbivorous lizards (Liolaemidae: Phymaturus) from the Payunia volcanic region of Argentina. Journal of Natural History 47(19-20): 1365-1378.

Corbalan, V.; Debandi, G. 2014. Resource segregation in two herbivorous species of mountain lizards from Argentina. Herpetological Journal 24(4): 201-208.

Lobo, F.; Abdala, C.S. 2007. Descripcion de una nueva especie de Phymaturus del grupo de P. palluma de la Provincia de Mendoza, Argentina. Cuadernos de Herpetologia 21(2): 103-113.

Phymaturus sinervoi

Scolaro, J.A.; Méndez-de la Cruz, F.R.; Ibargüengoytía, N.R. 2012. A new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) from isolated plateau of southwestern Rio Negro Province, Argentina. Zootaxa 3451: 17-30.

Phymaturus sitesi

Ávila, L.J.; Pérez, C.H.F.; Pérez, D.R.; Morando, M. 2011. Two new mountain lizard species of the Phymaturus genus (Squamata: Iguania) from northwestern Patagonia, Argentina. Zootaxa 2924: 1-21.

Phymaturus somuncurensis

Cruz, F.B.; Schulte, J.A.; Bellagamba, P. 1999. New distributional records and natural history notes for reptiles from southern Argentina. Herpetological Review 30(3): 182-183.

Phymaturus spectabilis

Boretto, J.M.; Cabezas-Cartes, F.; Kubisch, E.L.; Sinervo, B.; Ibargüengoytía, N.R. 2014. Changes in female reproduction and body condition in an endemic lizard, Phymaturus spectabilis, following the Puyehue volcanic ashfall event. Herpetological Conservation and Biology 9(1): 181-191.

Boretto, J.M.; Cabezas-Cartes, F.; Tappari, F.; Méndez-de la Cruz, F.R.; Sinervo, B.; Scolaro, A.J.; Ibargüengoytía, N.R. 2014. Reproductive biology of Phymaturus spectabilis (Liolaemidae): females skip reproduction in cold and harsh environments of Patagonia, Argentina. Herpetological Conservation and Biology 9(1): 170-180.

Cabezas-Cartes, F. 2013. Natural history notes: Phymaturus spectabilis. Predation. Herpetological Review 44(4): 679.

Cabezas-Cartes, F.; Leticia Kubisch, E.; Ibargüengoytía, N.R. 2014. Consequences of volcanic ash deposition on the locomotor performance of the Phymaturus spectabilis lizard from Patagonia, Argentina. Journal of Experimental Zoology Part A-Ecological Genetics and Physiology 321(3): 164-172.

Lobo, F.; Cruz, F.B.; Abdala, C.S. 2012. Multiple lines of evidence show that Phymaturus agilis Scolaro, Ibarguengoytia & Pincheira-Donoso, 2008 is a junior synonym of Phymaturus spectabilis Lobo & Quinteros, 2005. Cuadernos de Herpetologia 26(2): 21-27.

Lobo, F.; Quinteros, S. 2005. A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia (São Paulo) 45(13): 143-177.

Scolaro, J.A.; Ibargüengoytía, N.R.; Pincheira-Donoso, D. 2008. When starvation challenges the tradition of niche conservatism: on a new species of the saxicolous genus Phymaturus from Patagonia Argentina with pseudoarboreal foraging behaviour (Iguania, Liolaemidae). Zootaxa 1786: 48-60.

Phymaturus spurcus

Lobo, F.; Quinteros, S. 2005. Taxonomic studies of the genus Phymaturus (Iguania: Liolaemidae): Redescription of Phymaturus patagonicus Koslowsky 1898, and revalidation and redescription of Phymaturus spurcus Barbour 1921. Journal of Herpetology 39(4): 533-540.

Phymaturus tenebrosus

Kubisch, E.L.; Ippi, S.; Karlanian, J.; Ojeda, V. 2017. Distribution extension of a rare Phymaturus lizard (Squamata: Liolaemidae) reaching Nahuel Huapi National Park (Argentina). Herpetology Notes 10: 59-62.

Lobo, F.; Quinteros, S. 2005. A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia (São Paulo) 45(13): 143-177.

Phymaturus verdugo

Abdala, C.S.; Juarez, R. 2007. Geographic distribution: Phymaturus verdugo. Herpetological Review 38(1): 101-102.

Avila, L.J.; Fulvio Perez, C.H.; Perez, D.R.; Morando, M. 2007. Reptilia, Squamata, Liolaemidae, Phymaturus verdugo: distribution extension, new provincial records, filling gaps, and geographic distribution map. Check List 3(3): 250-252.

Cei, J.M.; Videla, F. 2003. A new Phymaturus species from volcanic Cordilleran mountains of the south-western Mendoza Province, Argentina (Liolaemidae, Iguania, Lacertilia, Reptilia). Museo Regionale di Scienze Naturali Bollettino (Turin) 20(2): 291-305.

Phymaturus videlai

Scolaro, J.A.; Pincheira-Donoso, D. 2010. Lizards at the end of the world: two new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) revealed in southern Patagonia of Argentina. Zootaxa 2393: 17-32.

Phymaturus vociferator

Pincheira-Donoso, D. 2004. Una nueva especie del genero Phymaturus (Iguania: Tropiduridae: Liolaemini) del centro-sur de Chile. Multequina 13: 57-70.

Urra, F.A.; Díaz, H.A.; Werning, H.; Eisenberg, T.; Troncoso-Palacios, J. 2017. Phymaturus vociferator Pincheira-Donoso, 2004 (Squamata: Liolaemidae): new records and updated geographic distribution. Check List 13(3)(Article 2137): 1-4.

Phymaturus williamsi

Fava, G.A.; Acosta, J.C.; Victorica, A.E.; Martínez, T.; Rodríguez, M. 2018. Natural history notes: Phymaturus williamsi. Predator-prey interaction. Herpetological Review 49(2): 331-332.

Lobo, F.; Laspiur, A.; Acosta, J.C. 2013. Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from northwestern Argentina. Zootaxa 3683(2): 117-132.

Phymaturus yachanana

Ávila, L.J.; Pérez, C.H.F.; Minoli, I.; Morando, M. 2014. A new lizard of the Phymaturus genus (Squamata: Liolaemidae) from Sierra Grande, northeastern Patagonia, Argentina. Zootaxa 3793(1): 99-118.

Phymaturus zapalensis

Boretto, J.M.; Ibargüengoytía, N.R. 2009. Phymaturus of Patagonia, Argentina: reproductive biology of Phymaturus zapalensis (Liolaemidae) and a comparison of sexual dimorphism within the genus. Journal of Herpetology 43(1): 96-104.

Boretto, J.M.; Ibargüengoytía, N.R. 2018. Natural history notes: Phymaturus zapalensis. Diet. Herpetological Review 49(4): 745.

Boretto, J.M.; Jahn, G.A.; Fornes, M.W.; Cussac, V.E.; Ibargüengoytía, N.R. 2012. How males synchronize their reproductive cycles with females to cope with seasonal climate: an endocrinal and ultrastructural study of Phymaturus zapalensis lizards (Liolaemidae). Herpetological Journal 22(1): 33-42.