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Taxonomy of the genus Protobothrops Biodiversity of the family Viperidae

Bibliography of the genus
Protobothrops (Habus and allies)

(Reptilia: Serpentes: Viperidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Protobothrops in general

Gumprecht, A.; Ryabov, S.A. 2002. Die Gattung Trimeresurus Lacepede, 1804 - Zum Kenntnisstand der Forschung. Draco 3(4) (12): 31-44.

Gumprecht, A.; Tillack, F.; Orlov, N.L.; Captain, A.; Ryabov, S. 2004. Asian Pitvipers. GeitjeBooks, Berlin. 368 pp.

Guo, P.; Liu, Q.; Wen, T.; Xiao, T.; Fang, M.; Zhong, G.; Nguyen, T.Q.; Zhu, F.; Jadin, R.C.; Li, C. 2016. Multilocus phylogeny of the Asian Lance-headed Pitvipers (Squamata, Viperidae, Protobothrops). Zootaxa 4093(3): 382-390.

Guo, P.; Pang, J.; Zhang, Y.P.; Zhao, E.M. 2006. A re-analysis of the phylogeny of the genus Protobothrops (Reptilia: Viperidae), with particular reference to the systematic position of P. xiangchengensis. Amphibia-Reptilia 27(3): 433-439.

Herrmann, H.W.; Ziegler, T.; Malhotra, A.; Thorpe, R.S.; Parkinson, C.L. 2004. Redescription and systematics of Trimeresurus cornutus (Serpentes: Viperidae) based on morphology and molecular data. Herpetologica 60(2): 211-221.

Hoge, A.R.; Romano-Hoge, S.A.R.W.L. 1983. Notes on micro and ultrastructure of 'Oberhautschen' in Viperoidea. Memorias do Instituto Butantan (Sao Paulo) 44-45 1980-1981[1983]: 81-118.

Huang, X.; Pan, T.; Han, D.; Zhang, L.; Hou, Y.; Yu, L.; Zheng, H.; Zhang, B. 2012. A new species of the genus Protobothrops (Squamata: Viperidae: Crotalinae) from the Dabie Mountains, Anhui, China. Asian Herpetological Research 3(3): 213-218.

Malhotra, A.; Thorpe, R.S. 2000. A phylogeny of the Trimeresurus group of pit vipers: new evidence from a mitochondrial gene tree. Molecular Phylogenetics and Evolution 16(2): 199-211.

Orlov, N.L.; Ryabov, S.A.; Shiryaev, K.A.; Bguyen, V.S. 2001. On the biology of pit vipers of Protobothrops genus (Serpentes: Colubroidea: Viperidae: Crotalinae). Russian Journal of Herpetology 8(2): 159-164.

Protobothrops cornutus

David, P.; Tong, H.; Vogel, G.; Tian, M. 2008. On the status of the Chinese pitviper Ceratrimeresurus shenlii Liang and Liu in Liang, 2003 (Serpentes, Viperidae) with the addition of Protobothrops cornutus (Smith, 1930) to the Chinese snake fauna. Asiatic Herpetological Research 11: 17-23.

Gong, S.P.; Hitschfeld, E.; Hundsdörfer, A.K.; Auer, M.; Wang, F.; Zhou, L.; Fritz, U. 2011. Is the horned pitviper Ceratrimeresurus shenlii Liang and Liu, 2003 from China a valid Protobothrops? Amphibia-Reptilia 32(1): 132-135.

Gong, S.P.; Wang, H.; Yang, C.T.; Yu, C.S.; Pan, D.S.; Wang, F. 2010. [Protobothrops cornutus discovered in Tianjingshan Forest, Guangdong Province, China]. (In Chinese). Chinese Journal of Zoology 45(3): 170-173.

Herrmann, H.W.; Ziegler, T.; Malhotra, A.; Thorpe, R.S.; Parkinson, C.L. 2004. Redescription and systematics of Trimeresurus cornutus (Serpentes: Viperidae) based on morphology and molecular data. Herpetologica 60(2): 211-221.

Le, K.Q.; Ziegler, T. 2009. Geographic distribution: Protobothrops cornutus (Smith's Horned Pit Viper). Vietnam: Ha Giang. Herpetological Review 40(1): 115.

Luu, V.Q.; Nguyen, T.Q.; Lehmann, T.; Bonkowski, M.; Ziegler, T. 2015. New records of the Horned Pitviper, Protobothrops cornutus (Smith, 1930) (Sepentes: Viperidae), from Vietnam with comments on morphological variation. Herpetology Notes 8: 149-152.

Orlov, N.L.; Ryabov, S.A.; Nguyen, V.S.; Nguyen, Q.T. 2003. New records and data on the poorly known snakes of Vietnam. Russian Journal of Herpetology 10(3): 217-240.

Shiryaev, K.A.; Orlov, N.L.; Ryabov, S.A.; Nguyen, Q.T. 2007. Captive breeding and reproductive biology of the Protobothrops cornutus and Triceratolepidophis sieversorum. Russian Journal of Herpetology 14(1): 57-64.

Smith, M.A. 1930. Two new snakes from Tonkin, Indo-China. Annals and Magazine of Natural History (Ser. 10) 6: 681-683.

Ziegler, T.; Herrmann, H.W. 2002. Weiderentdeckung zweier gehörnter Grubenotterarten in Vietnam. ZGAP Mitteilungen 18(2): 24-26.

Protobothrops dabieshanensis

Huang, X.; Pan, T.; Han, D.; Zhang, L.; Hou, Y.; Yu, L.; Zheng, H.; Zhang, B. 2012. A new species of the genus Protobothrops (Squamata: Viperidae: Crotalinae) from the Dabie Mountains, Anhui, China. Asian Herpetological Research 3(3): 213-218.

Zhang, B.; Huang, X.; Pan, T.; Zhang, L.; Zhou, W.; Song, T.; Han, D. 2013. Systematics and species validity of the Dabieshan Pit Viper Protobothrops dabieshanensis Huang et al. 2012: evidence from a mitochondrial gene sequence analysis. Asian Herpetological Research 4(4): 282-287.

Protobothrops elegans

Hokama, Z.; Kamura, T.; Nozaki, M. 1981. Study on experimental envenomation by Sakishima-habu (Trimeresurus elegans). The Snake 13(1): 27-31.

Kamura, T.; Nishimura, M. 1992. Two years survival of Trimeresurus elegans (Viperidae) without food consumption. The Snake 24(2): 165-166.

Keegan, H.L.; Yoshino, K. 1959. Trimeresurus elegans (Gray, 1849), the Takishima habu, a venomous pit viper from the Ryukyu Islands. American Journal of Tropical Medicine and Hygiene 8(2): 124-133.

Kikukawa, D. 1974. A venomous snake, Trimeresurus elegans. Collecting Breed. Tokyo 36(7): 152-156.

Neill, W.T. 1960. The caudal lure of various juvenile snakes. Quarterly Journal of the Florida Academy of Sciences 23: 173-200.

Nikai, T.; Komori, Y.; Suzuki, Y.; Sugihara, H. 1997. Methionyl-lysyl-bradykinin-releasing enzyme from the venom of Trimeresurus elegans. Journal of Natural Toxins 6(3): 243-259.

Nishimura, M.; Kamura, T. 2009. [Effects of incubation temperature on eggs and hatchlings of habu (Protobothrops flavoviridis: Viperidae) and Sakishimahabu (P. elegans)]. (In Japanese). Biological Magazine Okinawa 47: 41-45.

Terada, K. 2011. [The distribution, population density and controls of Protobothrops mucrosquamatus, Protobothrops elegans, Elaphe taeniura friesei, 3 snake species established on Okinawa Island]. (In Japanese). Bulletin of the Herpetological Society of Japan 2011(2): 161-168.

Protobothrops flavoviridis

Abe, Y.; Mishima, S. 1961. On the sexual characteristics and food habits of Habu (Trimeresurus flavoviridis) with bodily measurements. Jap. J. sanit. Zool. 12: 135.

Abe, Y.; Mori, T. 1960. Epidemiological studies on Habu-bite sickness in Amami Oshima. Jap. J. sanit. Zool. 11: 62-63.

Abe, Y.; Tanaka, H.; Mishima, S.; Ono, T. 1965. Studies on the habu (Trimeresurus flavoviridis) and the habu-bite in Amami Oshima (Tr. 3). Analytical studies on the locomotion of the habu with a special reference to the influences of illumination. Jap. J. sanit. Zool. 16: 177-183.

Araki, Y.; Nishimura, M. 1984. Notes on ecology of habu, Trimeresurus flavoviridis, based on the experience of Mr. T. Minami, a snake hunter. Biological Magazine Okinawa 22: 115-116.

Ford, N.B.; Seigel, R.A. 1989. Relationships among body size, clutch size, and egg size in three species of oviparous snakes. Herpetologica 45(1): 75-83.

Fukada, H. 1978. Effect of temperature on the heart beats and activity of the habu, Trimeresurus flavoviridis. Snake 10(1): 17-20, 94.

Fukada, H.; Kihara, H.; Hayashi, Y. 1979. Studies on attractants and repellents to habu using electrocardiogram. Snake 11(1): 16-19, 120.

Gloyd, H.K. 1955. A new Crotalid snake from Kume Shima, Riu Kiu Islands. Bulletin of the Chicago Academy of Sciences 10(8): 123-134.

Groves, F. 1959. Notes on Trimeresurus f. flavoviridis. Herpetologica 15: 182.

Hattori, S. 1999. Development of the box trap for habu. pp. 257-263. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Hattori, S. 2002. [Present state and problem of habu snake (Trimeresurus flavoviridis)]. (In Japanese, English summary). Occasional Papers Kagoshima University Research Center for the Pacific Islands 36: 15-21.

Hattori, S.; Noboru, Y.; Kihara, H.; Hayashi, Y. 1999. Trap capture of habu (Trimeresurus flavoviridis) with odor extracted from rats. pp. 264-267. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Hayashi, Y.; Hattori, S.; Tanaka, H. 1984. The electric net trap for the venomous snake, habu, Trimeresurus flavoviridis. Japanese Journal of Experimental Medicine 54(5): 207-210.

Hayashi, Y.; Kihara, H.; Tanaka, H. 1982. Immobilizing effect of venom of habu (Trimeresurus flavoviridis) on small rodents. The Snake 14(1): 35-39.

Hayashi, Y.; Kihara, H.; Tanaka, H.; Kurosawa, M. 1984. Evaluation of a bait trap for habu, the venomous snake, Trimeresurus flavoviridis. Japanese Journal of Experimental Medicine 54(4): 171-175.

Hayashi, Y.; Kihara, H.; Tanaka, H.; Noboru, Y.; Yamashita, H.; Minami, T. 1979. Studies on attractancy to habu (Trimeresurus flavoviridis) in the field of Tokunoshima Island. Snake 11(1): 45-53, 126-127.

Hayashi, Y.; Noboru, Y. 1979. Trapping survey on the number of the rats living indoors in Amamioshima and Tokunoshima. Snake 11(1): 63-66, 128.

Hayashi, Y.; Noboru, Y.; Kawaguchi, E.; Tanaka, H. 1978. Survey of frequency of observation of the habu by residents of the Setouchi-town. Snake 10(1): 59-64, 99.

Hayashi, Y.; Noboru, Y.; Tanaka, H. 1978. Prevention of invasion of the habu by an electric fence. Snake 10(1): 42-46, 98.

Hayashi, Y.; Noboru, Y.; Wada, Y.; Tanaka, H. 1979. The method to prevent the invasion of habu (Trimeresurus flavoviridis) by an electric fence. Snake 11(1): 58-62, 127-128.

Hayashi, Y.; Sawai, Y.; Tanaka, H.; Mishima, S. 1999. Development of electric fence barriers for habu (Trimeresurus flavoviridis) in the Amami Islands. pp. 319-326. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Hayashi, Y.; Tanaka, H. 1979. On the methods for the population estimation of habu (Trimeresurus flavoviridis). Snake 11(1): 72-77, 129.

Hayashi, Y.; Tanaka, H. 1981. Age determination in the venomous snake, habu, Trimeresurus flavoviridis. Japanese Journal of Experimental Medicine 51(4): 209-213.

Hayashi, Y.; Tanaka, H. 1982. Invasive behaviour of a venomous snake, habu, Trimeresurus flavoviridis, into residential area. Japanese Journal of Experimental Medicine 52(4): 209-211.

Hayashi, Y.; Tanaka, H.; Tanaka, H.; Natsushita, N.; Takai, R. 1979. Total body-length and sex composition of habu (Trimeresurus flavoviridis) in Tokunoshima Island. Snake 11(1): 67-72, 128-129.

Hayashi, Y.; Tanaka, H.; Wada, Y.; Mishima, S. 1983. The electric fence for preventing invasion of Trimeresurus flavoviridis, Habu, the venomous snake. Japanese Journal of Experimental Medicine 33(3): 165-172.

Hiragi, T.; Ikeuchi, E. 2015. [An observation of the predation on Okinawa Rail (Gallirallus okinawae) by Habu (Protobothrops flavoviridis)]. (In Japanese). Biological Magazine Okinawa 53: 45-47.

Hoge, A.R.; Romano-Hoge, S.A.R.W.L. 1983. Notes on micro and ultrastructure of 'Oberhautschen' in Viperoidea. Memorias do Instituto Butantan (Sao Paulo) 44-45 1980-1981[1983]: 81-118.

Hokama, Y.; Kondo, T.; Sawai, Y. 1988. Comparison of Trimeresurus flavoviridis venom in Amami-Oshima Tokunoshima and Okinawa Islands. The Snake 20(1): 20-24.

Hokama, Z. 1978. Study on experimental envenomation by the habu (Trimeresurus flavoviridis). Snake 10(2): 107-113.

Huntzinger, P.E. 2003. The Okinawa habu. Bulletin of the Chicago Herpetological Society 38(2): 25-28.

Ikeda, K.; Iwai, N.; Wada, Y.; Hayashi, Y. 1979. Technical development of the radiotracking system for studies of habu movement. Snake 11(1): 29-31, 122.

Ikeda, K.; Wada, Y.; Hayashi, Y.; Iwai, N.; Kihara, H.; Noboru, Y.; Yamashita, H. 1980. The wireless tracking of movement of habu, Trimeresurus flavoviridis. The Snake 10(1): 25-39, 96.

Ishii, A.; Sawai, Y. 1999. Biological control of habu with Entamoeba invadens. pp. 391-398. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Kardong, K.V. 1999. Venom delivery by the brown treesnake (Boiga irregularis) and the habu (Trimeresurus flavoviridis). pp. 128-138. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Kihara, H.; Hashimura, S.; Nakamoto, E.; Hayashi, Y. 1985. Sex composition and reproduction of habu, Trimeresurus flavoviridis, the venomous snake. The Snake 17(2): 135-139.

Kihara, H.; Hayashi, Y.; Wada, Y.; Yamashita, H.; Noboru, Y. 1979. Studies on assay methods of attractants to habu 2. The assay methods in the enclosure for habu. Snake 11(1): 54-58, 127.

Kihara, H.; Hayashi, Y.; Wakisaka, I. 1978. Studies on attractants to habu using traps. Snake 10(1): 46-55, 98.

Kihara, H.; Miyashita, M.; Wakisaka, I.; Hayashi, Y.; Tanaka, H.; Tsumita, T. 1978. Studies on assay methods of attractants to the habu. Snake 10(1): 6-10, 93.

Kihara, H.; Yamashita, H.; Hayashi, Y.; Wada, Y.; Tanaka, H. 1979. Development of assay methods of attractants to habu 1. The assay method in a laboratory. Snake 11(1): 11-15, 119.

Kikuchi, S.; Sawai, Y.; Toshioka, S.; Matsui, T.; Okuyama, Y. 1981. Electronmicroscopy of the scale of two Japanese snakes. 1. Trimeresurus flavoviridis flavoviridis and T. okinavensis. The Snake 13(1): 6-15.

Koba, K. 1963. Food of Trimeresurus flavoviridis flavoviridis and T. okinavensis in the Amami Group of the Loo Choo Islands. Mem. Fac. Educ. Kumamoto Univ. 11: 35-41.

Koba, K. 1964. Ecology of the habu, Trimeresurus flavoviridis flavoviridis. Jap. J. sanit. Zool. 15: 95.

Koba, K. 1971. Natural history of the habu, Trimeresurus flavoviridis (Hallowell). The Snake 3(2): 75-96.

Koba, K.; Morimoto, H.; Ono, T.; Nakamoto, E.; Tanaka, K.; Yoshizaki, K. 1973. Oviposition and hatching of the habu, Trimeresurus flavoviridis. Acta Herpetologica Japonica 4(1-4): 13.

Koba, K.; Nakamoto, E.; Ono, T.; Tanaka, K.; Baba, K. 1967. Oviposition and hatching of the habu, Trimeresurus flavoviridis in the Amami Islands, Japan. Mem. Fac. Educ. Kumamoto Univ. 15(1): 29-37.

Koba, K.; Yashizaki, K.; Nakamoto, E.; Ono, T.; Tanaka, K. 1973. Oviposition of the habu, Trimeresurus flavoviridis. Acta Herpetologica Japonica 4(1-4): 13.

Kurihara, T.; Hayashi, Y. 1978. A brief study on the response of habu, Trimeresurus flavoviridis, to ultrasonics. Snake 10(1): 23-24, 95.

Maeda, H. 1978. On the association of habu-bites with meteorological factors. Snake 10(1): 85-87, 102-103.

Mason, R.T. 1999. Integrated pest management: the case for pheromonal control of habu (Trimeresurus flavoviridis) and brown treesnakes (Boiga irregularis). pp. 196-205. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Minakami, K. 1979. An estimation of age and life-span of the genus Trimeresurus (Reptilia, Serpentes, Viperidae) on Amami Oshima Island, Japan. Journal of Herpetology 13(2): 147-152.

Minakami, K.; Nakamoto, E. 1980. Hatching and early development of habu, Trimeresurus flavoviridis. The Snake 12(1-2): 11-14.

Minakami, K.; Nakamoto, E.; Matsushita, N.; Toriire, Y.; Takehara, K.; Fukushima, H. 1980. Studies on body length composition of captured habu, Trimeresurus flavoviridis or Amami Oshima Island, Japan. The Snake 12(1-2): 8-10.

Minakami, K.; Ono, T. 1976. Body length composition of the habu Trimeresurus flavoviridis on Amami Oshima Island. Japanese Journal of Herpetology 6(3): 61-63.

Mishima, S. 1965. Sexual dimorphism not clear snake, Calliophis japonicus: Eggs and hatching of Habu, Tremeresurus flavoviridis, and food habit of the genus. Acta Herpetologica Japonica 1: 55-56.

Mishima, S. 1966. Studies on the poisonous snake 'habu', Trimeresurus flavoviridis flavoviridis. 1. Food habit of Trimeresurus flavoviridis flavoviridis on the Amami Islands. Jap. J. sanit. Zool. 17: 1-21.

Mishima, S. 1967. Studies on the poisonous snake 'habu', Trimeresurus flavoviridis, 3. Nocturnal activity of the 'habu' on Amami Oshima Island. Jap. J. sanit. Zool. 18: 27-31.

Mishima, S.; Hayashi, Y.; Sawai, Y. 1984. Effect of the improvement of the habitual environment on the population of habu, Trimeresurus flavoviridis. The Snake 16(2): 115-118.

Mishima, S.; Hayashi, Y.; Tanaka, H.; Sawai, Y. 1999. The effectiveness of habitat modifications for controlling habu populations on Tokunoshima. pp. 365-371. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Mishima, S.; Nakamoto, E.; Yonemaru, M.; Sawai, Y.; Yamamoto, H. 1978. Studies on control of the habus. (Trimeresurus flavoviridis) 1. Living conditions of the habus on the Yoro Island from data of captive snakes. Snake 10(2): 151-164.

Mishima, S.; Nakamoto, E.; Yonemaru, M.; Sawai, Y.; Yamamoto, H. 1979. Studies on control of the habus (Trimeresurus flavoviridis). 2. Living conditons of the habus on Yoro Island from data of captured snakes. 2. Snake 11(1): 134-142.

Mishima, S.; Sawai, Y. 1978. Living conditions of the habus on Tokunoshima Island from data of captive snakes. Japanese Journal of Herpetology 7(4): 108-109.

Mishima, S.; Sawai, Y. 1979. Study on the habitat of habu, Trimeresurus flavoviridis, at Tete and Todoroki villages in Tokunoshima. Snake 11(1): 113-116, 133.

Mishima, S.; Sawai, Y.; Chigusa, Y.; Yamamoto, H. 1984. Studies on control of habu (Trimeresurus flavoviridis) 3. An analysis of the effect of land improvement on the population of habus on the Tokunoshima Island by the information obtained through questionaire. The Snake 16(1): 63-70.

Mishima, S.; Tanaka, H.; Sawai, Y. 1999. The biology of the habu (Trimeresurus flavoviridis). pp. 29-43. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Mishima, S.; Toriba, M. 2010. [Measurements of body sizes of the habu on Amami-Oshima Island in 1960]. (In Japanese). Bulletin of the Herpetological Society of Japan 2010(2): 125-131.

Miyashita, M.; Wakisaka, I. 1979. A study on the background for the invasion of habu into the village at Ketokina, Tokunoshima. Snake 11(1): 109-112, 133.

Miyashita, M.; Wakisaka, I. 1979. Relationship between the invasion of habu into the premises and the environmental factors in Tokunoshima. Snake 11(1): 103-108, 132.

Nakamoto, E. 1990. A himehabu (Trimeresurus okinavensis) swallowing a habu (Trimeresurus flavoviridis). The Snake 22(2): 138-139.

Nakamoto, E.; Fukushima, H.; Sawai, Y. 1981. Longevity of habu (Trimeresurus flavoviridis) breeded in a hungry condition. The Snake 13(1): 61-62.

Nakamoto, E.; Sawai, Y. 1982. Observation on egg-laying in the Taiwan-habu, Trimeresurus mucrosquamatus (Cantor). The Snake 14(2): 145-146.

Nakamoto, E.; Toriba, M. 2008. [Largest record of a clutch size in Protobothrops flavoviridis]. (In Japanese, English summary). Bulletin of the Herpetological Society of Japan 2008(1): 14-15.

Nishimura, M. 1986. Deviations and changes in measurements of snakes, mainly habu, Trimeresurus flavoviridis, alive, dead and after preservation. Nanki Seibutu 28(2): 97-103.

Nishimura, M. 1986. Fluctuations of body weight of habu, Trimeresurus flavoviridis, under rearing conditions. The Snake 18(2): 92-100.

Nishimura, M. 1990. Age estimation of habu, Trimeresurus flavoviridis (Viperidae) using growth bands in angulares. Amphibia-Reptilia 11(4): 411-418.

Nishimura, M. 1993. Estimation of age and growth of habu, Trimeresurus flavoviridis (Serpentes: Viperidae), in the Okinawa Islands. Japanese Journal of Ecology (Sapporo) 43(2): 83-90.

Nishimura, M. 1993. Estimation of age structure of habu, Trimeresurus flavoviridis (Serpentes: Viperidae), in the middle and southern districts of Okinawa Island and Minna Island. Japanese Journal of Ecology (Sapporo) 43(3): 155-161.

Nishimura, M. 1995. Body mass loss in habu, Trimeresurus flavoviridis, without water supply in biotron. The Snake 27(1): 13-20.

Nishimura, M. 1998. Arrangement of oviductal eggs and intra-clutch egg shape modification in the habu, Trimeresurus flavoviridis (Viperidae). Herpetological Review 29(2): 76-77.

Nishimura, M. 1999. Structure and application of the slanting nylon-net fence to prevent dispersal of habu (Trimeresurus flavoviridis). pp. 313-318. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Nishimura, M. 2000. [Effects of climate factors on the activity of a viperid snake, habu (Trimeresurus flavoviridis) - based on the frequencies of bite cases of sleeping men in the Okinawa Islands]. (In Japanese, English summary). Japanese Journal of Ecology (Otsu) 50(2): 133-140.

Nishimura, M.; Akamine, H.; Takahira, K.; Hiraiwa, A.; Okudo, H. 1988. Frequencies of encountering habu, Trimeresurus flavoviridis, by the inhabitants in Okinawa Prefecture -the survey method and the estimated relative densities of habu in several towns and villages-. The Snake 20(2): 121-132.

Nishimura, M.; Araki, Y.; Ueda, H.; Kawashima, Y. 1991. Frequencies of prey items of habu, Trimeresurus flavoviridis (Viperidae), in the Okinawa Islands 1. The Snake 23(2): 81-83.

Nishimura, M.; Kamura, T. 1989. Variations in tail length of habu, Trimeresurus flavoviridis, in the Okinawa Islands. The Snake 21(1): 18-28.

Nishimura, M.; Kamura, T. 1992. Creamy white material ejected by male habu, Trimeresurus flavoviridis (Viperidae). Japanese Journal of Herpetology 14(4): 163-165.

Nishimura, M.; Kamura, T. 1993. Twin hatchlings of Trimeresurus flavoviridis (Viperidae). Herpetological Review 24(1): 14-15.

Nishimura, M.; Kamura, T. 1994. Body color polymorphism in the snake Trimeresurus flavoviridis on Okinawa Island, Japan. Herpetological Review 25(1): 8-9.

Nishimura, M.; Kamura, T. 1994. Body mass changes in Trimeresurus flavoviridis (Viperidae) on Okinawa Island, Japan. Researches on Population Ecology (Kyoto) 36(2): 173-179.

Nishimura, M.; Kamura, T. 1994. Survival rate and sex ratio in habu, Trimeresurus flavoviridis (Viperidae), on the subtropical Okinawa Island, Japan. Researches on Population Ecology (Kyoto) 36(1): 115-120.

Nishimura, M.; Kamura, T. 2001. [Oviposition date of a viperid snake, habu (Trimeresurus flavoviridis) - relating factors and the effects on the hatching date]. (In Japanese, English summary). Biological Magazine Okinawa 39: 11-17.

Nishimura, M.; Kamura, T. 2009. [Effects of incubation temperature on eggs and hatchlings of habu (Protobothrops flavoviridis: Viperidae) and Sakishimahabu (P. elegans)]. (In Japanese). Biological Magazine Okinawa 47: 41-45.

Nishimura, M.; Katsuren, S. 1994. Annual survival rate and age structure of habu, Trimeresurus flavoviridis (Viperidae), on Minna Island, Japan: estimation from removal trapping. Researches on Population Ecology (Kyoto) 36(2): 255-260.

Nobusaka, R.; Nakamoto, E.; Sawai, Y. 1994. Studies on the reproduction of habu (Trimeresurus flavoviridis) and coloration of the offsprings. The Snake 26(1): 1-9.

Ogawa, T.; Onoue, H.; Nakagawa, K.; Nomura, S.; Sueishi, K.; Hattori, S.; Kihara, H.; Ohno, M. 1995. Localization and expression of phospholipases A2 in Trimeresurus flavoviridis (Habu snake) venom gland. Toxicon 33(12): 1645-1652.

Rodda, G.H.; Sawai, Y.; Chiszar, D.; Tanaka, H. (eds.). 1999. Problem Snake Management: The Habu and the Brown Treesnake. Cornell University Press, Ithaka, N.Y. 534 pp.

Sawai, Y.; Fukuyama, T.; Kato, K.; Kawamura, Y.; Honma, M. 1967. Case report on hypersensitivity to habu (Trimeresurus flavoviridis) venom. Jap. J. Bact. 22: 58-60.

Sawai, Y.; Kawamura, Y. 1988. Habu bites on the Amami Islands of Japan in 1984. The Snake 20: 93-97.

Sawai, Y.; Kawamura, Y.; Araki, Y.; Tomihara, Y. 1999. A historical outlook on studies of habu (Trimeresurus flavoviridis) bites in the Amami and Okinawa Islands of Japan. pp. 107-115. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Sawai, Y.; Kawamura, Y.; Ebihara, I.; Okonogi, T.; Yamakawa, M. 1967. Studies on the improvement of treatment of habu (Trimeresurus flavoviridis) bites. 6. Habu bites on the Amami and Ryukyu Island in 1964. Jap. J. exp. Med. 37: 51-59.

Sawai, Y.; Makino, M.; Tateno, I.; Okonogi, T.; Mitsuhashi, S. 1962. Studies on the improvement of treatment of Habu snake (Trimeresurus flaroviridis) bite. 3. Clinical analysis and medical treatment of Habu snake bite on the Amami Islands. Jap. J. exp. Med. 32: 117-138.

Shiroma, H. 1984. A case of pseudo-hermaphroditism in habu, Trimeresurus flavoviridis. The Snake 16(1): 56-62.

Shiroma, H. 1989. On the population estimate of the habu, Trimeresurus flavoviridis, by removal method with traps. pp. 384-392. In: Matsui, M., Hikida, T. & Goris, R.C. (eds.). Current herpetology in East Asia. Proceedings of the second Japan-China Herpetological Symposium, Kyoto, July 1988. Herpetological Society of Japan, Kyoto. 521 pp.

Shiroma, H. 1998. Bio-assay of attractants for habu, Trimeresurus flavoviridis. The Snake 28(1-2): 65-67.

Shiroma, H. 2004. A bioassay of the attractive effect of mouse urine on the habu, Trimeresurus flavoviridis. Current Herpetology 23(1): 43-46.

Shiroma, H.; Akamine, H. 1999. Complete removal of habu (Trimeresurus flavoviridis) from a residential area by trapping. pp. 327-339. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Takara, T. 1954. [Trends in Habu [Trimeresurus flavoviridis] number and losses in Okinawa since the war]. Oyo Dobuts. Zasshi 18: 187-192.

Tanaka, H. 1973. Activity and behavior of habu, Trimeresurus flaviridis. The Snake 5(1-2): 116-132.

Tanaka, H.; Hayashi, Y.; Kihara, H.; Hattori, S.; Mishima, S.; Wada, Y.; Kurosawa, M.; Sawai, Y. 1987. Population control of habu, Trimeresurus flavoviridis, the venomous snake, studied on Tokunoshima Island by a research group fy 1980 to 1983. The Snake 19(1): 26-40.

Tanaka, H.; Hayashi, Y.; Wada, Y. 1979. Epidemiological studies on factors correlated with the snake bite by habu, Trimeresurus flavoviridis, on the Amami Islands. Snake 11(1): 79-83, 130-131.

Tanaka, H.; Hayashi, Y.; Wada, Y. 1999. Population density of habu on the Amami Islands, as estimated by removal methods. pp. 230-235. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Tanaka, H.; Mishima, S.; Ono, T. 1967. Relationship betwwen the activity of a viper, Trimeresurus flavoviridis (Hallowell, 1860) and the snake bite. Jap. J. sanit. Zool. 18: 113-118.

Tanaka, H.; Wada, Y.; Hayashi, Y.; Ikeda, K. 1999. Movements of habu, as observed by radio tracking in the field. pp. 224-229. In: Rodda, G.H.; Sawai, Y.; Chiszar, D. & Tanaka, H. (eds.). Problem snake management: the habu and the brown treesnake. Comstock, Ithaca & London. 534 pp.

Tanaka, H.; Wada, Y.; Oguma, Y.; Sasa, M. 1971. A method of population estimation for the habu, Trimeresurus flavoviridis. The Snake 3(1): 9-13.

Tateno, L.; Sawai, Y.; Kawamura, Y.; Okonogi, T. 1965. Studies on the improvement of treatment of the Habu snake (Trimeresurus flavoviridis) bite. 5. Factors influencing the prognosis of habu snake bite. Jap. J. exp. Med. 35: 85-98.

Teruya, K.; Nakachi, K. 1961. Observation on the relationship between weather condition and the incidence of Habu-bite in the Ryukyu Archipelago. Jap. J. sanit. Zool. 12: 136.

Toriba, M.; Hirabayashi, K. 1991. Fang replacement cycle in a habu, Trimeresurus flavoviridis. The Snake 23(2): 119-120.

Wada, Y.; Ikeda, K.; Kihara, H.; Hayashi, Y.; Iwai, N.; Noboru, Y.; Yamashita, H. 1979. Tracing of movement of habu, Trimeresurus flavoviridis, by radio tracking. Snake 11(1): 32-36, 123.

Wada, Y.; Kihara, H.; Hayashi, Y.; Tanaka, H. 1979. Areal distribution of habu density in Tokunoshima Is. estimated from the cases of habu bite in the premises. Snake 11(1): 37-43, 124.

Wada, Y.; Tanaka, H.; Ikeda, K.; Suzuki, H.; Motoi, E.; Noboru, Y.; Kawai, J.; Ono, T. 1972. Radio-tracking of movement of habu, Trimeresurus flavoviridis, 2. The Snake 4(1): 44-50.

Wada, Y.; Tanaka, H.; Oguma, Y.; Sasa, M.; Ikeda, K.; Motoi, E.; Kawai, J.; Ono, T. 1971. Radio-tracking of movement of habu, Trimeresurus flavoviridis. The Snake 3(1): 20-23.

Wakisaka, I.; Miyashita, M.; Ando, T.; Takano, A. 1978. Comparison of epidemiological features of habu-bites between Amamioshima and Tokunoshima. Snake 10(1): 76-85, 101-102.

Wakisaka, I.; Miyashita, M.; Ando, T.; Takano, A. 1978. Epidemiological features of habu-bites in Amami-islands. Snake 10(1): 66-75, 101.

Wakisaka, I.; Miyashita, M.; Ando, T.; Takano, A.; Kawabata, N.; Kihara, H. 1979. An epidemiological study of habu-bites on the Amami Islands. Snake 11(1): 83-102, 131-132.

Wakisaka, I.; Sawai, Y. 1978. Epidemiological studies for habu in Amami-islands. Snake 10(1): 65-90, 100-101.

Waters, R.M.; Chiszar, D.; Burghardt, G.M. 1996. Strike-induced chemosensory searching in the habu (Trimeresurus flavoviridis), an Asian pit viper. Journal of Herpetology 30(2): 147-151.

Yamagishi, H. 1974. Observations on the nocturnal activity of the habu with special reference to the intensity of illumination. The Snake 6(1): 37-43.

Yasuno, M. 1978. On the exterminatory measures for habu in Amami district. The problems from an ecological point of view. Snake 10(1): 90, 103.

Yokoyama, F.; Masumitu, M.; Yoshida, H. 1995. A method for distinction of sex of habu, Trimeresurus flavoviridis. The Snake 27(1): 61-63.

Yokoyama, F.; Yoshida, H. 1993. The reproductive cycle of the male habu, Trimeresurus flavoviridis. The Snake 25(1): 55-62.

Yokoyama, F.; Yoshida, H. 1994. The reproductive cycle of the female habu, Trimeresurus flavoviridis. Journal of Herpetology 28(1): 54-59.

Yoshida, C. 1989. Behavior of habus in choosing egg-laying cavities. pp. 472-481. In: Matsui, M., Hikida, T. & Goris, R.C. (eds.). Current herpetology in East Asia. Proceedings of the second Japan-China Herpetological Symposium, Kyoto, July 1988. Herpetological Society of Japan, Kyoto. 521 pp.

Protobothrops himalayanus

Pan, H.; Chettri, B.; Yang, D.; Jiang, K.; Wang, K.; Zhang, L.; Vogel, G. 2013. A new species of the genus Protobothrops (Squamata: Viperidae) from southern Tibet, China and Sikkim, India. Asian Herpetological Research 4(2): 109-115.

Protobothrops jerdonii

Burger, R.M. 2005. Natural history notes: Protobothrops jerdonii xanthomelas (Chinese Mountain Pitviper). Reproduction. Herpetological Review 36(3): 326.

Fan, L.S.; Guo, C.; Zhang, X. 2000. [Distribution and reproductive ecology of Protobothrops jerdonii]. (In Chinese). Sichuan Journal of Zoology 19(3): 185.

Guo, P. 2000. [Comparative studies on hemipenial morphology of three crotaline snakes]. (In Chinese). Chinese Journal of Zoology 35(6): 9-11.

Guo, P.; Liu, Q.; Cao, L.; Chen, X.; Jiang, K.; Wang, Y.; Malhotra, A. 2011. Molecular phylogeography of Jerdon's pitviper (Protobothrops jerdonii): importance of the uplift of the Tibetan plateau. Journal of Biogeography 38(12): 2326-2336.

Guo, P.; Lu, S.Q.; Huang, S.; Zhao, H.; Zhao, E.M. 2006. Hemipenial morphology of five Asian pitvipers, with a discussion on their taxonomy. Amphibia-Reptilia 27(1): 19-23.

Guo, P.; Malhotra, A.; Li, C.; Creer, S.; Pook, C.E.; Wen, T. 2009. Systematics of the Protobothrops jerdonii complex (Serpentes, Viperidae, Crotalinae) inferred from morphometric data and molecular phylogeny. Herpetological Journal 19(2): 85-96.

Hoge, A.R.; Romano-Hoge, S.A.R.W.L. 1983. Notes on micro and ultrastructure of 'Oberhautschen' in Viperoidea. Memorias do Instituto Butantan (Sao Paulo) 44-45 1980-1981[1983]: 81-118.

Liu, C.C. 1940. Life history of Trimeresurus jerdoni. Peking Nat. Hist. Bull. 14(4): 245-252.

Luo, J.; Wang, R.; Li, D.S.; Xiong, Y.L.; Zhang, Y.P. 1998. The karyotype of Protobothrops jerdonii. (In Chinese). Zoological Research 19(6): 495-496.

Orlov, N.L.; Ryabov, S.A.; Nguyen, V.S.; Nguyen, Q.T. 2003. New records and data on the poorly known snakes of Vietnam. Russian Journal of Herpetology 10(3): 217-240.

Orlov, N.L.; Ryabov, S.A.; Shiryaev, K.A.; Bguyen, V.S. 2001. On the biology of pit vipers of Protobothrops genus (Serpentes: Colubroidea: Viperidae: Crotalinae). Russian Journal of Herpetology 8(2): 159-164.

Wall, F. 1910. Notes on the viviparous habit of Jerdon's pit viper (Lachesis jerdoni) and observations of the foetal tooth in the unborn embryo. Journal of the Bombay Natural History Society 20: 231-234.

Ye, Y.; Deng, X. 1997. Five new records of snake in Hunan province. (In Chinese). Chinese Journal of Zoology 32(5): 46-47.

Zambre, A.; Sheth, C.; Dalvi, S.; Kulkarni, N. 2010. First record of Protobothrops jerdonii xanthomelas (Gunther, 1889) from Eaglenest Wildlife Sanctuary, India. Journal of the Bombay Natural History Society 106(2): 211-213.

Zeng, X.M.; Zhang, F. 2000. [The karyotypes and the primary taxonomical discussion on three species of the genus Trimeresurus sensu lato]. (In Chinese, English summary). Acta Zootaxonomica Sinica 25(3): 354-358.

Protobothrops kaulbacki

Bhide, K.S.; Captain, A.; Bhatt, B.B.; Gumprecht, A.; Tillack, F. 2008. Das Portrait - der Kurzbericht zum Titelbild: Protobothrops kaulbacki (Smith). Sauria (Berlin) 30(2): 1-2.

Guo, P.; Malhotra, A.; Li, P.P.; Pook, C.E.; Creer, S. 2007. New evidence on the phylogenetic position of the poorly known Asian pitviper Protobothrops kaulbacki (Serpentes: Viperidae: Crotalinae) with a redescription of the species and a revision of the genus Protobothrops. Herpetological Journal 17(4): 237-246.

Rao, D.Q.; Zhao, E.M. 2005. A new record from China - Protobothrops kaulbacki (Reptilia, Serpentes, Viperidae). Acta Zootaxonomica Sinica 30(1): 209-211.

Protobothrops maolanensis

Chen, T.B.; Luo, J.; Meng, Y.; Wen, B.H.; Jiang, K. 2013. [Discovery of Protobothrops maolanensis in Guangxi, with taxonomic discussion]. (In Chinese). Sichuan Journal of Zoology 32(1): 116-118.

Kropachev, I.I.; Shiryaev, K.A.; Nguyen, T.T.; Orlov, N.L. 2015. New record of Protobothrops cf. maolanensis in northeastern Vietnam, with data on its morphology and biology. Russian Journal of Herpetology 22(2): 93-102.

Liu, Q.; Myers, E.A.; Zhong, G.H.; Hu, J.; Zhao, H.; Guo, P. 2012. Molecular evidence on the systematic position of the lance-headed pitviper Protobothrops maolanensis Yang et al., 2011. Zootaxa 3178: 57-62.

Pham, A.V.; Pham, C.T.; Ngoc, V.H.; Ziegler, T.; Nguyen, T.Q. 2017. New records of amphibians and reptiles from Ha Giang Province, Vietnam. Herpetology Notes 10: 183-191.

Yang, J.H.; Orlov, N.L.; Wang, Y. 2011. A new species of pitviper of the genus Protobothrops from China (Squamata: Viperidae). Zootaxa 2936: 59-68.

Protobothrops mucrosquamatus

Chang, T.Y.; Mao, S.H.; Guo, Y.W. 1997. Cloning and expression of a cysteine-rich venom protein from Trimeresurus mucrosquamatus (Taiwan habu). Toxicon 35(6): 879-888.

Guo, P. 2000. [Comparative studies on hemipenial morphology of three crotaline snakes]. (In Chinese). Chinese Journal of Zoology 35(6): 9-11.

Guo, P.; Liu, Q.; Zhu, F.; Zhong, G.H.; Che, J.; Wang, P.; Xie, Y.L.; Murphy, R.W.; Malhotra, A. 2018. Multilocus phylogeography of the Brown-spotted Pitviper Protobothrops mucrosquamatus (Reptilia: Serpentes: Viperidae) sheds a new light on the diversification pattern in Asia. Molecular Phylogenetics and Evolution 133 [2019]: 82-91.

Guo, P.; Lu, S.Q.; Huang, S.; Zhao, H.; Zhao, E.M. 2006. Hemipenial morphology of five Asian pitvipers, with a discussion on their taxonomy. Amphibia-Reptilia 27(1): 19-23.

Guo, P.; Zhao, E.M. 2006. Comparison of skull morphology in nine Asian pit vipers (Serpentes: Crotalinae). Herpetological Journal 16(3): 305-313.

Hoge, A.R.; Romano-Hoge, S.A.R.W.L. 1983. Notes on micro and ultrastructure of 'Oberhautschen' in Viperoidea. Memorias do Instituto Butantan (Sao Paulo) 44-45 1980-1981[1983]: 81-118.

Huang, S.M.; Huang, S.P.; Chen, Y.H.; Tu, M.C. 2007. Thermal tolerance and altitudinal distribution of three Trimeresurus snakes (Viperidae: Crotalinae) in Taiwan. Zoological Studies 46(5): 592-599.

Lalremsanga, H.T.; Lalbiakzuala; Lalrinsanga 2017. Geographic distribution: Protobothrops mucrosquamatus (Brown Spotted Pitviper). Herpetological Review 48(1): 131.

Luu, V.Q.; Nguyen, T.Q.; Pham, C.T.; Dang, K.N.; Vu, T.N.; Miskovic, S.; Bonkowski, M.; Ziegler, T. 2013. No end in sight? Further new records of amphibians and reptiles from Phong Nha-Ke Bang National Park, Quang Binh Province, Vietnam. Biodiversity Journal 4(2): 285-300.

Mao, J.J.; Norval, G. 2006. Natural history notes: Protobothrops mucrosquamatus (Taiwanese Habu). Stomach contents. Herpetological Review 37(3): 353-354.

Mao, J.J.; Norval, G.; Kung, W.B.; Chang, H.N. 2006. Natural history notes: Bungarus multicinctus multicinctus (Many-banded Krait). Diet. Herpetological Review 37(3): 350.

Mao, S.H. 1993. Common terrestrial venomous snakes of Taiwan. National Museum of Natural Science Special Publications (Taichung) 5: 1-108.

Nguyen, Q.T.; Doan, V.K.; Ho, T.C.; Nguyen, V.S. 2007. Geographic distribution: Protobothrops mucrosquamatus (Brown-spotted Pitviper). Herpetological Review 38(1): 105.

Nguyen, T.Q.; Nguyen, T.V.; Pham, C.T.; Ong, A.V.; Ziegler, T. 2018. New records of snakes (Squamata: Serpentes) from Hoa Binh Province, northwestern Vietnam. Bonn Zoological Bulletin 67(1): 15–24.

Orlov, N.L.; Ryabov, S.A.; Shiryaev, K.A.; Bguyen, V.S. 2001. On the biology of pit vipers of Protobothrops genus (Serpentes: Colubroidea: Viperidae: Crotalinae). Russian Journal of Herpetology 8(2): 159-164.

Papenfuss, T.J. 2010. Protobothrops mucrosquamatus. The IUCN Red List of Threatened Species 2010: e.T178409A7540882. http://dx.doi.org/10.2305/IUCN.UK.2010-4.RLTS.T178409A7540882.en.

Prater, S.H. 1929. Note on a Formosan viper (Trimeresurus mucrosquamatus) from the North-East frontier. Journal of the Bombay Natural History Society 33: 998.

Sakia, J. 1929. Record of symptoms and treatment of a bite from a Formosan viper (Trimeresurus mucrosquamatus). Journal of the Bombay Natural History Society 33: 998.

Song, M. 1987. [Survey of the reptiles of southern Shaanxi]. (In Chinese, English summary). Acta Herpetologica Sinica 6(1): 59-64.

Terada, K. 2011. [The distribution, population density and controls of Protobothrops mucrosquamatus, Protobothrops elegans, Elaphe taeniura friesei, 3 snake species established on Okinawa Island]. (In Japanese). Bulletin of the Herpetological Society of Japan 2011(2): 161-168.

Vasaruchapong, T.; Laoungbua, P.; Tangrattanapibul, K.; Tawan, T.; Chanhome, L. 2017. Protobothrops mucrosquamatus (Cantor, 1839), a highly venomous species added to the snake fauna of Thailand (Squamata: Viperidae). Tropical Natural History 17(2): 111-115.

Vogel, G.; Brachtel, N. 2006. Zur Haltung von Protobothrops mucrosquamatus. pp. 21-32. In: Kunz, K. & Werning, H. 25. Terraristika Hamm. Natur und Tier-Verlag, Recklinghausen. 32 pp.

Wu, Y.; Shu, F.; Yang, Y.M.; Ju, W.B.; Guo, P. 2010. [A preliminary study on artificial reproduction of Protobothrops mucrosquamatus]. (In Chinese). Jingji Dongwu Xuebao 14(3): 143-145.

Zeng, X.M.; Zhang, F. 2000. [The karyotypes and the primary taxonomical discussion on three species of the genus Trimeresurus sensu lato]. (In Chinese, English summary). Acta Zootaxonomica Sinica 25(3): 354-358.

Zhong, G.; Liu, Q.; Li, C.; Peng, P.; Guo, P. 2017. Sexual dimorphism and geographic variation in the Asian Lance-headed Pitviper Protobothrops mucrosquamatus in the mainland China. Asian Herpetological Research 8(2): 118-122.

Ziegler, T.; Vu, N.T.; Le, K.Q.; Nguyen, Q.T.; Hallermann, J.; Thach, M.H.; Le, V.K. 2006. Neue Verbreitungsnachweise einiger wenig bekannter vietnamesischer Amphibien und Reptilien. Sauria (Berlin) 28(2): 29-40.

Protobothrops sieversorum

Herrmann, H.W.; Ziegler, T.; Stuart, B.L.; Orlov, N.L. 2003. New findings on the distribution, morphology, and natural history of Triceratolepidophis sieversorum (Serpentes: Viperidae). Herpetological Natural History 9(1): 89-94.

Orlov, N.L.; Ryabov, S.A.; Nguyen, V.S.; Nguyen, Q.T. 2003. New records and data on the poorly known snakes of Vietnam. Russian Journal of Herpetology 10(3): 217-240.

Shiryaev, K.A.; Orlov, N.L.; Ryabov, S.A.; Nguyen, Q.T. 2007. Captive breeding and reproductive biology of the Protobothrops cornutus and Triceratolepidophis sieversorum. Russian Journal of Herpetology 14(1): 57-64.

Ziegler, T.; Herrmann, H.W. 2002. Weiderentdeckung zweier gehörnter Grubenotterarten in Vietnam. ZGAP Mitteilungen 18(2): 24-26.

Ziegler, T.; Herrmann, H.W.; David, P.; Orlov, N.L.; Pauwels, O.S.G. 2000. Triceratolepidophis sieversorum, a new genus and species of pitviper (Reptilia: Serpentes: Viperidae: Crotalinae) from Vietnam. Russian Journal of Herpetology 7(3): 199-214.

Protobothrops tokarensis

Koba, K.; Kikukawa, D. 1970. A taxonomic study of the Tokarahabu, Trimeresurus tokarensis from the Tokara Islands, Japan. Bull. biogeogr. Soc. Japan 25: 1-8.

Koba, K.; Kikukawa, D. 1971. A taxonomic study of the Tokara-habu, Trimeresurus tokarensis Nagai. The Snake 3(1): 39-52.

Koba, K.; Kikukawa, D. 1971. Notes on the tokara-habu Trimeresurus tokarensis from Kodakara-jima. Acta Herpetologica Japonica 4(1-4): 12.

Koba, K.; Tanaka, K.; Nakamoto, E.; Morimoto, H. 1973. Oviposition of the tokara-habu, Trimeresurus tokarensis. Acta Herpetologica Japonica 4(1-4): 13.

Protobothrops trungkhanhensis

Orlov, N.L.; Ryabov, S.A.; Nguyen, T.T. 2009. Two new species of genera Protobothrops Hoge et Romano-Hoge, 1983 and Viridovipera Malhotra et Thorpe, 2004 (Ophidia: Viperidae: Crotalinae) from Karst region in northeastern Vietnam. Part I. Description of a new species of Protobothrops genus. Russian Journal of Herpetology 16(1): 69-82.

Protobothrops xiangchengensis

David, P.; Tong, H. 1997. Translations of recent descriptions of Chinese pitvipers of the Trimeresurus-complex (serpentes, viperidae), with a key to the complex in China and adjacent areas. Smithsonian Herpetological Information Service 112: 1-31.

Guo, P.; Malhotra, A.; Li, C.; Creer, S.; Pook, C.E.; Wen, T. 2009. Systematics of the Protobothrops jerdonii complex (Serpentes, Viperidae, Crotalinae) inferred from morphometric data and molecular phylogeny. Herpetological Journal 19(2): 85-96.

Guo, P.; Zhang, F. 2001. Comparative studies on hemipenes of four species of Trimeresurus (sensu stricto) (Serpentes: Crotalinae). Amphibia-Reptilia 22(1): 113-117.

Guo, P.; Zhao, E.M. 2006. Comparison of skull morphology in nine Asian pit vipers (Serpentes: Crotalinae). Herpetological Journal 16(3): 305-313.