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Taxonomy of the genus Ptyodactylus Biodiversity of the family Phyllodactylidae

Bibliography of the genus Ptyodactylus (Fan-toed Geckos)

(Reptilia: Sauria: Phyllodactylidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Ptyodactylus in general

Böhme, W. 2007. Editorial linguistic deficiency causes a superfluous nomen nudum in the gekkonid genus Ptyodactylus (Squamata: Gekkonidae). Herpetological Review 38(1): 12.

Boutan, L. 1893. Memoire sur les reptiles rapportes de Syrie par Theodore Barrois: 1, genre Ptyodactyle. Rev. Biol. Nord France 5: 329-345, 368-384, 444-448.

Frankenberg, E. 1974. Vocalization of males of three geographical forms of Ptyodactylus from Israel (Reptilia: Sauria: Gekkonidae). Journal of Herpetology 8(1): 59-70.

Frankenberg, E. 1976. Daily activity of geographical forms of Ptyodactylus (Reptilia: Gekkonidae). Israel Journal of Zoology 25(4): 207.

Frankenberg, E.; Werner, Y.L. 1980. Effect of lunar cycle on daily activity rhythm in a gekkonid lizard, Ptyodactylus. Israel Journal of Zoology 28(4): 244-228.

Gamble, T.; Bauer, A.M.; Greenbaum, E.; Jackman, T.R. 2008. Out of the blue: a novel, trans-Atlantic clade of geckos (Gekkota, Squamata). Zoologica Scripta 37(4): 355-366.

Heimes, P. 1987. Beitrag zur Systematik der Fächerfinger (Sauria: Gekkonidae: Ptyodactylus). Salamandra 23(4): 212-235.

Lymberakis, P.; Kaliontzopoulou, A. 2003. Additions to the herpetofauna of Syria. Zoology in the Middle East 29: 33-39.

Metallinou, M.; Cervenka, J.; Crochet, P.A.; Kratochvil, L.; Wilms, T.; Geniez, P.; Shobrak, M.Y.; Brito, J.C.; Carranza, S. 2015. Species on the rocks: systematics and biogeography of the rock-dwelling Ptyodactylus geckos (Squamata: Phyllodactylidae) in North Africa and Arabia. Molecular Phylogenetics and Evolution 85: 208-220.

Sallam, F.A.E. 2005. Genetic variation in populations of the fan-fingered gecko "Ptyodactylus" from Egypt: electrophoretic evidence. Journal of Union of Arab Biologists Cairo A Zoology 24: 215-231.

Werner, Y.L. 1965. Über die israelischen Geckos der Gattung Ptyodactylus und ihre Biologie. Salamandra 1: 15-25.

Werner, Y.L. 2006. Retraction of Ptyodactylus Goldfuss from the fauna of Iran and its replacement by a new species of Asaccus Dixon and Anderson (Reptilia: Sauria: Gekkonidae). Hamadryad 30(1-2): 135-140.

Werner, Y.L.; Chernis, E.; Granot, R. 2008. The call of the Sri Lankan golden gecko Calodactylodes illingworthorum, ecological parallel of the Fan-toed Geckos, genus Ptyodactylus (Reptilia: Sauria: Gekkonidae). Bioacoustics 18(1): 35-49.

Werner, Y.L.; Frankenberg, E. 1982. Multidimensional investigation of a group of animals, exemplified by Ptyodactylus (Reptilia: Sauria: Gekkonidae). pp. 60-63. In: Barth, F.G. Neurobiology and strategies of adaption Symposium, 1981, Goethe Universität, Frankfurt.

Werner, Y.L.; Goldenzweig, A. 2018. Natural history notes: Ptyodactylus spp. (Fan-toed Geckos). Field body temperatures. Herpetological Review 49(2): 332.

Werner, Y.L.; Sivan, N. 1993. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the Levant: 1. Biometry of three species in Israel. Revista Espanola de Herpetologia 7: 47-64.

Werner, Y.L.; Sivan, N. 1994. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the Levant: 2, taxonomy, with a review of ecology and zoogeography. Revista Espanola de Herpetologia 8: 105-122.

Ptyodactylus ananjevae

Nazarov, R.A.; Melnikov, D.A.; Melnikova, E. 2013. Three new species of Ptyodactylus (Reptilia; Squamata; Phyllodactylidae) from the Middle East. Russian Journal of Herpetology 20(2): 147-162.

Ptyodactylus dhofarensis

Nazarov, R.A.; Melnikov, D.A.; Melnikova, E. 2013. Three new species of Ptyodactylus (Reptilia; Squamata; Phyllodactylidae) from the Middle East. Russian Journal of Herpetology 20(2): 147-162.

Ptyodactylus guttatus

Ali, R.A.M. 2012. Genetic variation among nine Egyptian gecko species (Reptilia: Gekkonidae) based on RAPD-PCR. Life Science Journal Acta Zhengzhou University Overseas Edition 9(1): 154-162.

Arad, Z. 1995. Physiological responses to increasing ambient temperature in three ecologically different, congeneric lizards (Gekkoninae: Ptyodactylus). Comparative Biochemistry and Physiology A Comparative Physiology A 112(2): 305-311.

Arad, Z.; Raber, P.; Werner, Y.L. 1989. Selected body temperature in diurnal and nocturnal forms of Ptyodactylus (Reptilia: Gekkoninae) in a photothermal gradient. Journal of Herpetology 23(2): 103-108.

Blondheim, S.A.; Frankenberg, E. 1983. 'Protest' sounds of a grasshopper: predator-deterrent signal? Psyche (Cambridge) 90(4): 387-394.

Blondheim, S.A.; Frankenberg, E. 1984. Adaptive significance of protest sounds in a grasshopper. (Abstract). Israel Journal of Zoology 32(2-3): 156.

Boutan, L. 1892. Excursion zoologique a la Montagne de Hummoun ul Faroun (Montagne des bains de Pharaon). Arch. Z. exper. 2(10): 1-22.

Frankenberg, E. 1979. Influence of light and temperature on daily activity patterns of three Israeli forms of Ptyodactylus (Reptilia: Gekkoninae). Journal of Zoology (London) 189(1): 21-30.

Frankenberg, E. 1984. Meet your subtenant - the house gecko. Israel Land and Nature 9(4): 149-155.

Goldberg, S.R. 2011. Natural history notes: Ptyodactylus guttatus (Sinai Fan-fingered Gecko). Reproduction. Herpetological Review 42(3): 433.

Johnston, G.; Bouskila, A. 2007. Sexual dimorphism and ecology of the gecko, Ptyodactylus guttatus. Journal of Herpetology 41(3): 506-513.

Mienis, H.K. 1987. A snail in the stomach of the gecko Ptyodactylus hasselquistii guttatus. Kreukel (Amsterdam) 23(7): 107-108.

Nagy, K.A.; Degen, A.A. 1988. Do desert geckos conserve energy and water by being nocturnal? Physiological Zoology 61(6): 495-499.

Perry, G.; Brandeis, M. 1992. Variation in stomach contents of the gecko Ptyodactylus hasselquistii guttatus in relation to sex, age, season and locality. Amphibia-Reptilia 13(3): 275-282.

Rosenberg, H.I.; Russell, A.P.; Cavey, M.J. 1992. Development of the subdigital adhesive pads of Ptyodactylus guttatus (Reptilia: Gekkonidae). Journal of Morphology 211(3): 243-258.

Werner, D. 1973. Beobachtungen an Ptyodactylus hasselquistiiguttatus (Geckonidae). Verhandlungen der Naturforschenden Gesellschaft in Basel 82(1): 54-87.

Werner, Y.L. 1986. Ecology of eggs and laying sites of Ptyodactylus geckos. pp. 441-444. In: Rocek, Z. (ed.). Studies in herpetology. Proceedings of the European Herpetological Meeting (3rd Ordinary General Meeting of the Societas Europaea Herpetologica) Prague 1985. Charles University, Prague. 754 pp.

Werner, Y.L. 1995. Some unusual accidental herpetological finds from Cyprus and Lebanon, including a new Ptyodactylus (Reptilia: Lacertilia: Gekkonidae). Biologia Gallo-Hellenica 22: 67-76.

Werner, Y.L.; Rothenstein, D.; Sivan, N. 1991. Directional asymmetry in reptiles (Sauria: Gekkonidae: Ptyodactylus) and its possible evolutionary role, with implications for biometrical methodology. Journal of Zoology (London) 225(4): 647-658.

Werner, Y.L.; Sivan, N. 1996. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the Levant: 3, experimental and natural hybrids of P. guttatus and P. puiseuxi. Israel Journal of Zoology 42(2): 185-202.

Zlotkin, E.; Milman, T.; Sion, G.; Werner, Y.L. 2003. Predatory behaviour of gekkonid lizards, Ptyodactylus spp., towards the scorpion Leiurus quinquestriatus hebraeus, and their tolerance of its venom. Journal of Natural History 37(5): 641-646.

Ptyodactylus hasselquistii

Ali, R.A.M. 2012. Genetic variation among nine Egyptian gecko species (Reptilia: Gekkonidae) based on RAPD-PCR. Life Science Journal Acta Zhengzhou University Overseas Edition 9(1): 154-162.

Al-Shammari, A.M. 2012. Additional records of lizards in Ha'il Province, Saudi Arabia. Russian Journal of Herpetology 19(4): 287-291.

Angel, F. 1950. Contribution a l'étude de l'Air (Mission L. Chopard et A. Villiers). Lezards. Mem. Inst. Franc. Afr. Noire 10: 331-336.

Arad, Z. 1995. Physiological responses to increasing ambient temperature in three ecologically different, congeneric lizards (Gekkoninae: Ptyodactylus). Comparative Biochemistry and Physiology A Comparative Physiology A 112(2): 305-311.

Arad, Z.; Raber, P.; Werner, Y.L. 1989. Selected body temperature in diurnal and nocturnal forms of Ptyodactylus (Reptilia: Gekkoninae) in a photothermal gradient. Journal of Herpetology 23(2): 103-108.

Baha el Din, S.M. 1999. On the specific status of Ptyodactylus ragazzii Anderson, 1898 and its occurrence in Egypt (Squamata: Sauria: Gekkonidae). Herpetozoa 12(1-2): 39-43.

Baha el Din, S.M.; Attum, O. 2000. The herpetofauna of Zaranik Protected Area, Egypt, with notes on their ecology and conservation. Herpetological Bulletin 73: 17-21.

Boulenger, G.A. 1896. A list of the reptiles and batrachians collected by the late Prince Eugenio Ruspoli in Somaliland and Gallaland in 1893. Annali del Museo Civico de Storia Naturale di Genova (Ser. 2) 17: 5-16.

Boutan, L. 1892. Excursion zoologique a la Montagne de Hummoun ul Faroun (Montagne des bains de Pharaon). Arch. Z. exper. 2(10): 1-22.

Carrillo de Espinoza, N.; Rothenstein, D.; Salas, A.; Werner, Y.L. 1990. Radiation and convergence among desert geckos: Phyllodactylus species resembling both Ptyodactylus and Stenodactylus. Amphibia-Reptilia 11(1): 1-13.

Cunningham, P.L. 2008. Ptyodactylus hasselquistii (Donndorff, 1789). Egyptian Fan-footed Gecko. Termite Diet. African Herp News 45: 6-7.

El Ghazaly, N.; Dekinesh, S.I.; Moursi, A.A.; Hussein, H.K.; Abdou, H.A.M. 1995. Studies on the growth rates of three lizard species under the influence of normal and artificial conditions of light and temperature. Journal of the Egyptian German Society of Zoology (A) 18: 413-434.

El Ghazaly, N.A.; Hussein, H.K. 1991. Thermal behaviour, heat exchange and lethal temperature in two lizard species, Agama stellio and Ptychodactylus hasselquistii. Journal of the Egyptian German Society of Zoology 5: 165-176.

Esterbauer, H. 1990. Ökologische und verhaltensbiologische Betrachtungen am Fächerfingergecko. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 43(1): 26-30.

Frankenberg, E. 1979. Influence of light and temperature on daily activity patterns of three Israeli forms of Ptyodactylus (Reptilia: Gekkoninae). Journal of Zoology (London) 189(1): 21-30.

Frankenberg, E. 1984. Meet your subtenant - the house gecko. Israel Land and Nature 9(4): 149-155.

Fritz, J.P.; Schutte, F. 1987. Geckos der Gattungen Ptyodactylus und Hemidactylus aus der Arabischen Republik Jemen. Bonner Zoologische Beiträge 38(2): 115-128.

Grandison, A.G.C. 1956. On a collection of lizards from West Africa. Bulletin de l'Institut Fondamental d'Afrique Noire Serie A Sciences Naturelles 18: 224-245.

Haas, G. 1957. Some amphibians and reptiles from Arabia. Proceedings of the California Academy of Sciences (Ser. 4) 29(4): 47-86.

Haas, G.; Battersby, J.C. 1959. Amphibians and reptiles from Arabia. Copeia 1959: 196-202.

Haupl, M. 1980. Das Schädelskelett einiger Arten der Fam. Gekkonidae. Annalen des Naturhistorischen Museums in Wien 83: 479-518.

Hussein, K.H. 1993. Seasonal variation of daily activity times in gekkonid lizards inhabiting Burg El-Arab region, west of Alexandria. Journal of the Egyptian German Society of Zoology (A) 11: 231-245.

Ibrahim, A.A. 2001. Geographic distribution: Ptyodactylus hasselquistii hasselquistii (Fan-toed Gecko). Herpetological Review 32(2): 120.

Ibrahim, A.A. 2002. The reptile community in the Zaranik Protected Area, northern Sinai, Egypt with special reference to their ecology and conservation. Journal of Union of Arab Biologists Cairo A Zoology 17: 1-16.

Mienis, H.K. 1986. A bicephalic Ptyodactylus hasselquisti from Israel. Zoology in the Middle East 1: 90-92.

Mohammed, M.B.H. 1989. Some observations on the cartilaginous structure of the head skeleton in some geckos (Reptilia: Gekkonidae). Zoologischer Anzeiger (Leipzig) 222(1-2): 3-11.

Rickert, M. 1995. Die Fächerfingergeckos der Gattung Ptyodactylus. Teil 1: Haltung und Zucht von Ptyodactylus hasselquistii hasselquistii (Donndorff, 1798) - Vergleichsmerkmale der nordafrikanischen und arabischen Form. Sauria (Berlin) 17(4): 19-28.

Rösler, H. 2000. Studien an den Begattungsorganen der Geckos (Reptilia: Gekkonomorpha) - 4. Der Hemipenis von Ptyodactylus hasselquistii hasselquistii (Donndorff, 1798) (Gekkonidae: Gekkonidae). Gekkota 2: 249-254.

Scortecci, G. 1940. Biologia del Tibesti. Ann. Mus. libico Stor. nat. 2: 257-275.

Werner, Y.L. 1964. Frequencies of regenerated tails, and structure of caudal vertebrae in Israeli desert geckos (Reptilia: Gekkonidae). Israel Journal of Zoology 13: 134-136.

Werner, Y.L. 1968. Regeneration frequencies in geckos of two ecologiccal types (Reptilia: Gekkonidae). Vie et Milieu (C) 19: 199-221.

Werner, Y.L. 1995. Some unusual accidental herpetological finds from Cyprus and Lebanon, including a new Ptyodactylus (Reptilia: Lacertilia: Gekkonidae). Biologia Gallo-Hellenica 22: 67-76.

Werner, Y.L.; Frankenberg, E.; Adar, O. 1978. Further observations on the distinctive vocal repertoire of Ptyodactylus hasselquistii cf. hasselquistii (Reptilia: Gekkoninae). Israel Journal of Zoology 27(4): 176-188.

Werner, Y.L.; Goldblatt, A. 1978. Body temperature in a basking gekkonid lizard, Ptyodactylus hasselquistii (Reptilia, Lacertilia, Gekkonidae). Journal of Herpetology 12(3): 408-411.

Werner, Y.L.; Rothenstein, D.; Sivan, N. 1991. Directional asymmetry in reptiles (Sauria: Gekkonidae: Ptyodactylus) and its possible evolutionary role, with implications for biometrical methodology. Journal of Zoology (London) 225(4): 647-658.

Zari, T.A. 1999. Seasonal acclimatization in metabolic rate of the fan-fingered gecko, Ptyodactylus hasselquistii (Reptilia: Gekkonidae). Journal of Thermal Biology 24(2): 137-142.

Ptyodactylus orlovi

Nazarov, R.A.; Melnikov, D.A.; Melnikova, E. 2013. Three new species of Ptyodactylus (Reptilia; Squamata; Phyllodactylidae) from the Middle East. Russian Journal of Herpetology 20(2): 147-162.

Zwanzig, B.M. 2018. Data on the vertical distribution and ecology of Ptyodactylus orlovi Nazarov, Melnikov & Melnikova, 2013. Sauria (Berlin) 40(2): 75 - 78.

Ptyodactylus oudrii

Garcia-Marquez, M.; Lopez-Jurado, L.F.; Mateo, J.A. 1996. Puestas comunales en el geco Ptyodactylus oudrii. Boletin de la Asociacion Herpetologica Espanola 7: 28-30.

Lataste, F. 1880. Descriptions of new species of reptiles from Algiers. Le Nat. 1880: 299, 306, 325.

Mateo, J.A.; Cuadrado, M. 1998. Natural history notes: Ptyodactylus oudrii (NCN). Parental care. Herpetological Review 29(2): 101-102.

Mateo, J.A.; Cuadrado, M. 2012. Communal nesting and parental care in Oudri's Fan-footed Gecko (Ptyodactylus oudrii): field and experimental evidence of an adaptive behavior. Journal of Herpetology 46(2): 209-212.

Perera, A.; Harris, D.J. 2010. Genetic variability within the Oudri's fan-footed gecko Ptyodactylus oudrii in north Africa assessed using mitochondrial and nuclear DNA sequences. Molecular Phylogenetics and Evolution 54(2): 634-639.

Scheler, B. 2013. Zur Haltung und Vermehrung des Fächerfingergeckos Ptyodactylus oudrii Lataste, 1880. Sauria (Berlin) 35(4): 3-13.

Ptyodactylus puiseuxi

Afrasiab, S.R.; Mohamad, S.I. 2009. A study on cave-dwelling geckos in Iraq, with the description of a new species from Saffine mountain (Reptilia: Gekkonidae). Zoology in the Middle East 47: 49-56.

Arad, Z. 1995. Physiological responses to increasing ambient temperature in three ecologically different, congeneric lizards (Gekkoninae: Ptyodactylus). Comparative Biochemistry and Physiology A Comparative Physiology A 112(2): 305-311.

Arad, Z.; Raber, P.; Werner, Y.L. 1989. Selected body temperature in diurnal and nocturnal forms of Ptyodactylus (Reptilia: Gekkoninae) in a photothermal gradient. Journal of Herpetology 23(2): 103-108.

Frankenberg, E. 1979. Influence of light and temperature on daily activity patterns of three Israeli forms of Ptyodactylus (Reptilia: Gekkoninae). Journal of Zoology (London) 189(1): 21-30.

Frankenberg, E. 1984. Meet your subtenant - the house gecko. Israel Land and Nature 9(4): 149-155.

Goldberg, S.R.; Maza, E. 2017. Natural history notes: Ptyodactylus puiseuxi (Israeli Fan-fingered Gecko). Reproduction. Herpetological Review 48(4): 853-854.

Werner, Y.L.; Rothenstein, D.; Sivan, N. 1991. Directional asymmetry in reptiles (Sauria: Gekkonidae: Ptyodactylus) and its possible evolutionary role, with implications for biometrical methodology. Journal of Zoology (London) 225(4): 647-658.

Werner, Y.L.; Sivan, N. 1996. Systematics and zoogeography of Ptyodactylus (Reptilia: Sauria: Gekkonidae) in the Levant: 3, experimental and natural hybrids of P. guttatus and P. puiseuxi. Israel Journal of Zoology 42(2): 185-202.

Wever, E.G.; Hepp-Reymond, M.C. 1967. Auditory sensitivity in the fan-toed gecko Pytodactylus hasselquistii puiseuxi Boutan. Proceedings of the National Academy of Sciences of the United States of America 57: 681-687.

Zlotkin, E.; Milman, T.; Sion, G.; Werner, Y.L. 2003. Predatory behaviour of gekkonid lizards, Ptyodactylus spp., towards the scorpion Leiurus quinquestriatus hebraeus, and their tolerance of its venom. Journal of Natural History 37(5): 641-646.

Ptyodactylus ragazzii

Baha el Din, S.M. 1999. On the specific status of Ptyodactylus ragazzii Anderson, 1898 and its occurrence in Egypt (Squamata: Sauria: Gekkonidae). Herpetozoa 12(1-2): 39-43.

Rickert, M. 1998. Die Fächerfingergeckos der Gattung Ptyodactylus. Teil 2: Haltung und Zucht von Ptyodactylus hasselquistii ragazzi Anderson, 1898. Sauria (Berlin) 20(2): 3-9.

Riva, I. de la; Padial, J.M. 2008. First record of the genus Ptyodactylus Goldfuss, 1820 (Sauria: Gekkonidae) for Mauritania (West Africa). Salamandra 44(1): 51-53.

Seufer, H. 1988. Zur Haltung und Zucht des Fächerfingergeckos Ptyodactylus hasselquistii ragazzi Anderson, 1898. Herpetofauna (Weinstadt) 10(55): 6-12.

Ptyodactylus rivapadialorum

Trape, J.F. 2017. Une nouvelle espèce de gecko palmé du genre Ptyodactylus Goldfuss, 1820 (Sauria, Ptyodactilidae) de Mauritanie. Bulletin de la Societe Herpetologique de France 163: 53-66.

Ptyodactylus ruusaljibalicus

Simó-Riudalbas, M.; Metallinou, M.; Pous, P. de; Els, J.; Jayasinghe, S.; Péntek-Zakar, E.; Wilms, T.; Al-Saadi, S.; Carranza, S. 2017. Cryptic diversity in Ptyodactylus (Reptilia: Gekkonidae) from the northern Hajar Mountains of Oman and the United Arab Emirates uncovered by an integrative taxonomic approach. PLoS ONE 12(8): e0180397.