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Taxonomy of the genus Timon Biodiversity of the family Lacertidae

Bibliography of the genus Timon (Ocellated Lizards)

(Reptilia: Sauria: Lacertidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Timon in general

Ahmadzadeh, F.; Carretero, M.A.; James, H.D.; Perera, A.; Böhme, W. 2012. A molecular phylogeny of the eastern group of ocellated lizard genus Timon (Sauria: Lacertidae) based on mitochondrial and nuclear DNA sequences. Amphibia-Reptilia 33(1): 1-10.

Ahmadzadeh, F.; Flecks, M.; Carretero, M.A.; Böhme, W.; Ihlow, F.; Kapli, P.; Miraldo, A.; Rödder, D. 2016. Separate histories in both sides of the Mediterranean: phylogeny and niche evolution of ocellated lizards. Journal of Biogeography 43(6): 1242-1253.

Arets, M.H.M. 2017. Europäische Perleidechsen: Arten und Unterarten der Gattung Timon. L@certidae 2017(6): 168-182.

Arnold, E.N.; Arribas, O.J.; Carranza, S. 2007. Systematics of the Palaearctic and Oriental lizard tribe Lacertini (Squamata: Lacertidae: Lacertinae), with descriptions of eight new genera. Zootaxa 1430: 3-86.

Bischoff, W. 1993. Übersicht der Arten und Unterarten der Familie Lacertidae. 1. Nachtrag. Die Eidechse 4(9): 15-16.

Corti, C.; Bischoff, W.; Böhme, W. 1999. Die Lacertiden der Mittelmeer-Inseln. Ein erste Übersicht. Die Eidechse 10(2): 41-58.

Joger, U. 1981. Zur Herpetofaunistik Westafrikas. Bonner Zoologische Beiträge 32(3-4): 297-340.

Mateo, J.A.; Lopez-Jurado, L.F.; Guillaume, C.P. 1996. Variabilite electrophoretique et morphologique des lézards ocelles (Lacertidae): un complexe d'especes de part et d'autre du detroit de Gibraltar. Comptes Rendus de l'Academie des Sciences Serie III Sciences de la Vie A 319(8): 737-746.

Mayer, W.; Bischoff, W. 1996. Beiträge zur taxonomischen Revision der Gattung Lacerta (Reptilia: Lacertidae). Teil 1: Zootoca, Omanosaura, Timon und Teira als eigenständige Gattungen. Salamandra 32(3): 163-170.

Timon kurdistanicus

Lazic, M.M.; Rödder, D.; Kaliontzopoulou, A. 2017. The ontogeny of developmental buffering in lizard head shape. Evolution & Development 19(6): 244-252

Rykena, S.; Bischoff, W. 1997. Kurzporträt: Timon princeps kurdistanicus (Suchov, 1936). Kurdische Zagros-Eidechse. Die Eidechse 8(2): 41-43.

Rykena, S.; Nettmann, H.K. 1986. The karyotype of Lacerta princeps kurdistanica and its meaning in phylogeny. pp. 193-196. In: Rocek, Z. (ed.). Studies in herpetology. Proceedings of the European Herpetological Meeting (3rd Ordinary General Meeting of the Societas Europaea Herpetologica) Prague 1985. Charles University, Prague. 754 pp.

Rykena, S.; Nettmann, H.K.; Bings, W. 1977. Zur Biologie der Zagros-Eidechse, Lacerta princeps Blanford 1874. 1. Beobachtungen im Freiland und im Terrarium an Lacerta princeps kurdistanica Suchov 1936 (Reptilia: Sauria: Lacertidae). Salamandra 13(3-4): 174-184.

Timon lepidus

Anonymous. 2001. Lacerta lepida. Jewelled lizard, ocellated lizard, Daudin, 1802. Reptilia (GB) 16: 39-42.

Allen, A. 1977. Changes in population density of the eyed lizard, Lacerta lepida, at three localities in Portugal between 1969 and 1975. British Journal of Herpetology 5(9): 661-662.

Arribas, O.J.; Cliville, S. 1994. Albinismo en Lacerta lepida Daudin, 1802 (Reptilia: Lacertidae). Boletin de la Asociacion Herpetologica Espanola 5: 20-23.

Bania, S. 2002. Eine der schönsten Echsen Europas im Terrarium: die Perleidechse. Reptilia (D) 7(37): 71-74.

Bannert, H. 1994. Bericht über Freiland-Haltung und erfolgreiche Nachzucht im Freien von Lacerta lepida in Österreich. Die Eidechse 5(13): 8-9.

Baranowitz, S.A.; Maderson, P.F.A.; Connelly, T.G. 1979. Lizard and newt tail regeneration: a quantitative study. Journal of Experimental Zoology 210(1): 17-38.

Bastin, E. 1983. Observations herpetologiques effectuees durant le sejour des Naturalistes belges en Andorre, du 4 au 14 juillet 1981. Naturalistes Belges 64(5): 163-172.

Berroneau, M. 2013. Mise á jour de la faune herpétologique du département du Lot-et-Garonne. Bulletin de la Societe Herpetologique de France 144: 23-32.

Bischoff, W. 1982. Zur Frage der taxonomischen Stellung europäischer und nordwestafrikanischer Perleidechsen (Sauria, Lacertidae, Lacerta lepida - Gruppe). Amphibia-Reptilia 2(4): 357-367.

Bischoff, W. 1985. Lacerta (Timon) lepida Daudun. Sauria (Berlin) 7(3): 19-24.

Bischoff, W.; Cheylan, M.; Böhme, W. 1984. Lacerta lepida - Perleidechse. pp. 181-210. In: Böhme, W. (ed.). Handbuch der Reptilien und Amphibien Europas. Band 2/1. Echsen (Sauria) 2. (Lacertidae 2: Lacerta). AULA-Verlag, Wiesbaden. 416 pp.

Böhme, W.; Esser, S. 2015. A remarkable age and size record of a male Jewelled Lizard, Timon lepidus (Daudin, 1802). Herpetozoa 28(1-2): 104-106.

Brugiere, D. 1987. Le lezard ocelle (Lacerta lepida) dans le departement du Cantal. Bulletin de la Societe Herpetologique de France 44: 6-7.

Busack, S.D.; Visnaw, J.A. 1989. Observations on the natural history of Lacerta lepida in Cadiz Province, Spain. Amphibia-Reptilia 10(3): 201-213.

Bustard, R. 1959. Lizards from southern Europe. Aquarist and Pondkeeper 24: 70-71.

Cabanas, R. 1946. Nota sobre un caso de hiperregeneracion de la cola de (Lacerta ocellata). Bol. Soc. esp. Hist, nat. 48(9-10): 525-526.

Cancio, I.; Gonzalez-Robles, A.; Bastida, J.M.; Manzaneda, A.J.; Salido, T.; Rey, P.J. 2016. Habitat loss exacerbates regional extinction risk of the keystone semiarid shrub Ziziphus lotus through collapsing the seed dispersal service by foxes (Vulpes vulpes). Biodiversity and Conservation 25(4): 693-709.

Castilla, A.M.; Bauwens, D. 1989. Reproductive characteristics of the lacertid lizard Lacerta lepida. Amphibia-Reptilia 10(4): 445-452.

Castilla, A.M.; Bauwens, D. 1990. Reproductive and fat body cycles of the lizard, Lacerta lepida, in central Spain. Journal of Herpetology 24(3): 261-266.

Castilla, A.M.; Bauwens, D. 1992. Habitat selection by the lizard Lacerta lepida in a Mediterranean oak forest. Herpetological Journal 2(1): 27-30.

Castilla, A.M.; Bauwens, D.; Llorente, G.A. 1991. Diet composition of the lizard Lacerta lepida in central Spain. Journal of Herpetology 25(1): 30-36.

Castilla, M.A.; Castanet, J. 1986. Growth, age and longevity of Lacerta lepida assessed by skeletochronology. pp. 331-335. In: Rocek, Z. (ed.). Studies in herpetology. Proceedings of the European Herpetological Meeting (3rd Ordinary General Meeting of the Societas Europaea Herpetologica) Prague 1985. Charles University, Prague. 754 pp.

Castilla, M.A.; Mateo, J.A. 1987. Comparative study of the reproduction of Lacerta lepida (Reptilia: Lacertidae) in different regions of Spain. pp. 91-94. In: van Gelder, J.J.; Strijbosch, H. & Bergers, P.J.M. (eds.). Proceedings of the 4th Ordinary General Meeting of the Societas Europaea Herpetologica. Societas Europaea Herpetologica, Faculty of Sciences, Nijmegen. 473 pp.

Ceia-Hasse, A.; Sinervo, B.; Vicente, L.; Pereira, H.M. 2014. Integrating ecophysiological models into species distribution projections of European reptile range shifts in response to climate change. Ecography 37(7): 679-688.

Chaignon, H. de 1905. Notes zoologiques. Autun, Bul. soc. hist. nat. 18: 150-157.

Cheylan, M. 1984. Croissance et determination de l'age chez le lezard ocelle (groupe Lacerta lepida, Sauria, Lacertidae) de France et du Maroc a partir de la squelettochronologie. Bulletin du Museum d'Histoire Naturelle de Marseille 44: 29-37.

Cheylan, M.; Grillet, P. 2004. Le lezard ocelle. Belin Eveil Nature, Paris. 95 pp.

Cheylan, M.; Grillet, P. 2005. Statut passe et actuel du lezard ocelle (Lacerta lepida, sauriens, lacertides) en France. Implication en termes de conservation. Vie et Milieu 55(1): 15-30.

Decaux, C. 1896. Observationa sur un Lezard ocellé en captivité depuis treize ans. C. R. Congr. Soc. Sav. Sorbonne 1896: 192-203.

Deeming, D.C. 1988. Eggshell structure of lizards of two sub-families of the Gekkonidae. Herpetological Journal 1(6): 230-234.

Delibes, M.; Palacios, F.; Garzon, J.; Castroviejo, J. 1975. Notes sur l'alimentation et la biologie du lynx pardelle, Lynx pardina (Temminck, 1824), en Espagne. Mammalia 39(3): 387-393.

Desclos, M. 1995. Note sur les circonstances curieuses, de la capture d'un jeune lezard ocelle Lacerta lepida (Daudin, 1802) dan le departement des Landes. Bulletin de la Societe Herpetologique de France 69-70: 69.

Deso, G.; Dusoulier, F.; Bence, S.; Cheylan, M. 2011. Distribution du Lézard Ocellé Timon lepidus lepidus (Daudin, 1802) dans la Haute vallée de la Durance (Région Provence-Alpes-Côte d’Azur). Bulletin de la Societe Herpetologique de France 137: 43-50.

Diaz, J.A.; Monasterio, C.; Salvador, A. 2006. Abundance, microhabitat selection and conservation of eyed lizards (Lacerta lepida): a radiotelemetric study. Journal of Zoology (London) 268(3): 295-301.

Díaz-Ruiz, F.; Ferreras, P. 2011. [Predation on Timon lepidus by a feral cat]. (In Spanish). Boletin de la Asociacion Herpetologica Espanola 22: 148-151.

Domingo Delgado, J.; Gomez, M. de los A. 2016. Evidencia del uso de drenaje viario por Timon lepidus (Daudin, 1802) en Andalucia occidental. Anales de Biologia (Murcia) 38: 63-67

Domingo, M.A. 1995. Nuevos datos sobre la distribucion del lagarto ocelado Lacerta lepida (Daudin, 1802) en la provincia de Alava (Pais Vasco). Munibe (Ciencias Naturales Natur Zientziak) 47: 111-112.

Dore, F.; Cheylan, M.; Grillet, P. 2015. Le Lézard ocellé - Un géant sur le continent européen. Biotope, Mèze. 191 pp.

Dore, F.; Grillet, P.; Thirion, J.M.; Besnard, A.; Cheylan, M. 2011. [A long-term monitoring program of the Ocellated Lizard (Timon lepidus) population of Oleron Island]. (In French). Revue d'Ecologie la Terre et la Vie 66(3): 205-214.

Dore, F.; Grillet, P.; Thirion, J.M.; Besnard, A.; Cheylan, M. 2011. Implementation of a long-term monitoring program of the ocellated lizard (Timon lepidus) population on Oleron Island. Amphibia-Reptilia 32(2): 159-166.

Duhaldeborde, F. 2012. Les zones militaires comme acteurs de la protection des espèces: dans le site landais de DGA Essais de Missiles (Biscarrosse), des biotopes refuges pour deux plantes et un reptile remarquables. Bulletin de la Societe Linneenne de Bordeaux 40(2): 151-158.

Esser, S.; Böhme, W. 2009. (Vorläufiger) Altersrekord einer Perleidechse, Timon lepidus (Daudin, 1802), im Tierhaus des Zoologischen Forschungsmuseums Alexander Koenig in Bonn. Elaphe 17(4): 44-47.

Ewald, P. 1989. Lacerta lepida. pp. 130-131. In: Castanet, J. & Guyetant, R. Atlas de repartition des amphibiens et reptiles de France. Societe Herpetologique de France, Paris. 191 pp.

Fei, T.; Skidmore, A.K.; Valentijn, V.; Wang, T.; Schlerf, M.; Toxopeus, B.; Overjijk, S. van; Bian, M.; Liu, Y.L. 2012. A body temperature model for lizards as estimated from the thermal environment. Journal of Thermal Biology 37(1): 56-64.

Fei, T.; Skidmore, A.K.; Valentijn, V.; Wang, T.; Toxopeus, B.; Bian, M.; Liu, Y.L. 2012. Predicting micro thermal habitat of lizards in a dynamic thermal environment. Ecological Modelling 231: 126-133.

Ferrand de Almeida, F.; Ferrand de Almeida, N.M.; Ferrand de Almeida, P. 1985. Nova contribuicao para o conhecimento da fauna do Paul de Arzilla. Ciencia Biologica Ecology and Systematics 5(2-3): 191-194.

Ferreira, D.; Mateus, C.; Santos, X. 2016. Responses of reptiles to fire in transition zones are mediated by bioregion affinity of species. Biodiversity and Conservation 25(8): 1543-1557.

Ferri, V.; dell'Acqua, A.; Salvidio, S. 1991. Distribuzione dei rettili nella fascia costiera della Liguria occidentale: 1. Lacerta l. lepida e Malpolon m. monspessulanus. Supplemento alle Ricerche di Biologia della Selvaggina 16: 217-220.

Font, E.; Lanuza, G.P. i de; Sampedro, C. 2009. Ultraviolet reflectance and cryptic sexual dichromatism in the ocellated lizard, Lacerta (Timon) lepida (Squamata: Lacertidae). Biological Journal of the Linnean Society 97(4): 766-780.

Frisch, O. von 1969. Europas größte Echse. Perleidechsen in Südfrankreich. Aquarien Magazin 3: 381-383.

Frisch, O. von 1969. Europas grösste Echse. Perleidechsen in Südfrankreich. Aquarien Magazin 3: 381-383.

Frisch, O. von 1980. Über ökologische Veranderungen und neue faunistische Beobachtungen in der Crau (Südfrankreich) 1969-1980. Bonner Zoologische Beiträge 31(3-4): 199-206.

Funke, O. 1999. Standorttreue bei Timon lepidus lepidus. Die Eidechse 10(2): 63.

Galan, P. 2000. Conducta gregaria del lagarto ocelado, Lacerta lepida, en las Islas Cies (Galicia). Boletin de la Asociacion Herpetologica Espanola 11(2): 80-82.

García-Díaz, P.; Ayres, C. 2011. Natural history notes: Timon lepidus (Ocellated Lizard). Predation. Herpetological Review 42(1): 96.

Ghiglione, C.; Crovetto, F.; Maggesi, M.; Maffei, S. 2017. Use of an artificial refuge for oviposition by a female Ocellated Lizard (Timon lepidus) in Italy. Herpetological Bulletin 136: 29-30.

Gil Sanchez, J.M. 1998. Seleccion de presa por el aguila-azor perdicera (Hieraaetus fasciatus) durante el periodo de nidificacion en la provincia de Granada (SE de Espana). Ardeola 45(2): 151-160.

Grillet, P.; Cheylan, M.; Dusoulier, F. 2006. Evolution des habitats et changement climatique: quelles consequences pour les populations de lezard ocelle, Lacerta lepida (Saurien, Lacertides), en limite nord de repartition? Ecologia Mediterranea 32: 63-72.

Grillet, P.; Cheylan, M.; Thirion, J.M.; Dore, F.; Bonnet, X.; Dauge, C.; Chollet, S.; Marchand, M.A. 2010. Rabbit burrows or artificial refuges are a critical habitat component for the threatened lizard, Timon lepidus (Sauria, Lacertidae). Biodiversity and Conservation 19(7): 2039-2051.

Grillet, P.; Thirion, J.M. 1999. Redecouverte de la repartition du lezard ocelle Lacerta lepida Daudin, 1802 en Charente-Maritime. Annales de la Societe des Sciences Naturelles de la Charente-maritime 8(8): 941-945.

Grillet, P.; Thirion, J.M.; Cheylan, M. 2010. Caracterisation de l'activite alimentaire annuelle du lezard ocelle Timon lepidus a partir des feces sur l'ile d'Oleron (Littoral Atlantique francais). Revue d'Ecologie la Terre et la Vie 65(3): 255-264.

Hahne, A. 1994. Urlaubsbekanntschaften mit Perleidechsen in Portugal. Die Eidechse 5(11): 11-12.

Hahne, A.; Fenske, R. 1992. Haltung und Zucht von Lacerta lepida. Erfahrungen aus Terrarien- und Freilandhaltung. Die Eidechse 3(6): 3-10.

Hahne, A.; Fenske, R. 1993. Haltung und Zucht von Lacerta l. lepida Daudin, 1802 - Erfahrungen aus Terrarien- und Freilandhaltung. Herpetofauna (Weinstadt) 15(82): 9-14.

Hahne, A.; Fenske, R. 1994. Haltung und Zucht der Perleidechse Lacerta lepida lepida Daudin, 1802 - Erfahrungen aus Terrarien- und Freilandhaltung. Sauria (Berlin) 16(3): 17-23.

Helmdag, A. 1992. Beobachtung einer ungewöhnlichen Paarung zwischen Gallotia stehlini und Lacerta lepida. Die Eidechse 3(6): 18.

Heredia Armada, B. 1983. Alimentacion invernal del elanio azul (Elanus caeruleus) en la provincia de Salamanca. Alytes (Merida) 1: 255-265.

Hernandez, A. 1990. Observaciones sobre el papel del lagarto ocelado (Lacerta lepida Daudin), el erizo (Erinaceus europaeus L.) y el tejon (Meles meles L.) en la dispersion desemillas. Doñana Acta Vertebrata 17(2): 235-242.

Hernandez, A.; Alegre, J.; Salgado, J.M. 1991. Ecologia trofica de Lacerta lepida en la provincia de Leon, noroeste de Espana. Amphibia-Reptilia 12(3): 283-292.

Hernandez, A.; Alegre, J.; Salgado, J.M.; Gutierrez, A. 1992. El papel de los coleopteros en el regimen alimentario de algunos vertebrados en el noroeste de Espana. Elytron (Barcelona) 5: 231-237.

Hodar, J.A.; Campos, F.; Rosales, B.A. 1996. Trophic ecology of the Ocellated Lizard Lacerta lepida in an arid zone of southern Spain: relationships with availability and daily activity of prey. Journal of Arid Environments 33(1): 95-107.

Hoffmann, E.G. 1963. Reproduction and raising of jewel lizards, Lacerta lepida lepida. Bulletin of the Philadelphia Herpetological Society 11(3-4): 53-55.

Howard, C.J. 1980. Eye lizards and True-lite. Herptile 5(3): 8-9.

Howard, C.J. 1985. Notes on breeding the eyed lizard (Lacerta lepida). Herptile 10(1): 18-23.

Howard, C.J. 2001. Notes on breeding the eyed lizard (Lacerta lepida). Herptile 26(1): 41-45.

Isenbugel, E.; Frank, W. 1987. Legenot bei einer Lacerta lepida Daudin, 1802 und einem Chamaeleo chamaeleon (Linnaeus, 1758) - eine Methode zur Austreibung von Resteiern. Salamandra 23(2-3): 179-180.

Juana, E. de; Juana, F. de 1982. Observacion de Lacerta lepida depredando un nido de Alectoris rufa. Doñana Acta Vertebrata 9: 374-375.

Kneller, M. 2011. Timon lepidus (Daudin, 1802) auf der portugiesischen Ilha da Berlenga? Die Eidechse 22(1): 25-26.

Kober, I. 2004. Die Perleidechse: Timon lepidus. Natur und Tier-Verlag, Münster. 62 pp.

Kober, I. 2013. Freilandhaltung von Perleidechsen. Reptilia (D) 18(101): 22-33.

Koci, P. 1967. [Observations on a case of reproduction of Lacerta lepida Daudin in captivity]. (In Polish, English summary). Przeglad Zoologiczny 11: 441-444.

Kroniger, M. 1996. Lloret de Mar - Einige kurze herpetologische Notizen. Die Eidechse 7(18): 31-32.

Laferrere, M. 1970. Observations erpetologiques. Riviera Scient. 1970: 89-90.

Lantermann, W.; Lantermann, Y. 2008. Perleidechsen im Freiland und im Terrarium. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 61(11): 68-73.

Lantermann, W.; Lantermann, Y. 2013. Perleidechsen (Timon lepidus) in der Sierra de Grazalema, Andalusien. Terraria-Elaphe 2013(42): 42-45.

Lanza, B.; Cei, J.M.; Crespo, E.G. 1977. Immunological investigations on the taxonomic status of some Mediterranean lizards (Reptilia Lacertidae). Monitore Zoologico Italiano New Series 11(3-4): 211-221.

Laplana Conesa, C.; Cuenca Bescos, G. 1995. Los microvertebrados (anfibios, reptiles y mamiferos) asociados al yacimiento de la Edad del Bronce de La Balsa la Tamariz (Tauste, Zaragoza). Col-Pa (Madrid) 47: 55-69.

Lataste, F. 1930. Presentation d'un lezard ocelle provenant de l'Herbe (Bassin d'Arcachon). Actes de la Société Linnéenne de Bordeaux 82: 69.

Lutz, D.; Mayer, W. 1984. Albumin-immunologische und proteinelektrophoretische Untersuchungen zur systematischen Stellung von Lacerta lepida Daudin und Lacerta princeps Blanford (Sauria, Lacertidae). Zoologischer Anzeiger (Leipzig) 212(1-2): 95-104.

Martin, J.; Lopez, P. 1996. Avian predation on a large lizard (Lacerta lepida) found at low population densities in Mediterranean habitats: an analysis of bird diets. Copeia 1996(3): 722-726.

Martin, J.; Lopez, P. 2002. The effect of Mediterranean dehesa management on lizard distribution and conservation. Biological Conservation 108(2): 213-219.

Martin, J.; López, P. 2010. Multimodal sexual signals in male ocellated lizards Lacerta lepida: vitamin E in scent and green coloration may signal male quality in different sensory channels. Naturwissenschaften 97(6): 545-553.

Martinez de Castilla, A. 1985. El lagarto ocelado. Vida Silvestre 53: 50-59.

Marty, P. 1900. Le Lezard ocelle (Lacerta ocellata Daud.) dans le Cantal. Feuille Natural 31: 27-28.

Mateo, J.A.; Castanet, J. 1994. Reproductive strategies in three Spanish populations of the Ocellated Lizard, Lacerta lepida (Sauria, Lacertidae). Acta Oecologica 15(2): 215-229.

Mateo, J.A.; Castroviejo, J. 1991. Variation morphologique et revision taxonomique de l'espece Lacerta lepida Daudin, 1802 (Sauria, Lacertidae). Bulletin du Museum National d'Histoire Naturelle Section A Zoologie Biologie et Ecologie Animales 12(3-4): 691-706.

Mateo, J.A.; Lopez-Jurado, L.F. 1997. Dental ontogeny in Lacerta lepida (Sauria, Lacertidae) and its relationship to diet. Copeia 1997(2): 461-463.

Mateo, J.A.; Lopez-Jurado, L.F.; Cano, J.; Guillaume, C.P. 1999. Intraspecific variability in nucleolar organiser position in Lacerta (Timon) lepida. Amphibia-Reptilia 20(1): 73-77.

Matos, A. de 2000. Meddelelse om et langtlevende perlefirben, Lacerta lepida = Timon lepidus. [Account of an aged ocellated lizard, Lacerta lepida = Timon lepidus]. (In Danish, English summary). Nordisk Herpetologisk Forening 43(5): 148-152.

Melani, E. 2006. Timon lepidus. Keeping and breeding the ocellated lizard. Reptilia (GB) 44: 60-65.

Mertens, R.; Wermuth, H. 1964. Über Lacerta margaritata Schinz, 1833, als ein 'nomen substitutum'. Bonner Zoologische Beiträge 15(1-2): 114.

Michels, J.P.; Bauer, A.M. 2004. Some corrections to the scientific names of amphibians and reptiles. Bonner Zoologische Beiträge 52(1-2): 83-94.

Miraldo, A.; Faria, C.; Hewitt, G.M.; Paulo, O.S.; Emerson, B.C. 2013. Genetic analysis of a contact zone between two lineages of the ocellated lizard (Lacerta lepida Daudin 1802) in south-eastern Iberia reveal a steep and narrow hybrid zone. Journal of Zoological Systematics and Evolutionary Research 51(1): 45-54.

Miraldo, A.; Hewitt, G.M.; Dear, P.H.; Paulo, O.S.; Emerson, B.C. 2012. Numts help to reconstruct the demographic history of the ocellated lizard (Lacerta lepida) in a secondary contact zone. Molecular Ecology 21(4): 1005-1018.

Miraldo, A.; Hewitt, G.M.; Paulo, O.S.; Emerson, B.C. 2011. Phylogeography and demographic history of Lacerta lepida in the Iberian Peninsula: multiple refugia, range expansions and secondary contact zones. BMC Evolutionary Biology 11 (Article 170): 19 pp.

Müller, L. 1939. Über eine auffällige Mutation des Farbenkleides bei einer jungen Lacerta lepida lepida Daudin. Zoologischer Anzeiger 128: 91-97.

Nicolau, S.M. 1956. Anfibios y reptiles de la Pobla de Segur. Pinneos 12(39-42): 45-55.

Nunes, V.L.; Beaumont, M.A.; Butlin, R.K.; Paulo, O.S. 2011. Multiple approaches to detect outliers in a genome scan for selection in ocellated lizards (Lacerta lepida) along an environmental gradient. Molecular Ecology 20(2): 193-205.

Odierna, G.; Olmo, E.; Capriglione, T.; Caputo, V. 1990. Karyological differences between Lacerta lepida and Lacerta pater. Journal of Herpetology 24(1): 97-99.

Ortega-Rubio, A. 1991. Trophic partitioning and community organization in a guild of lizards in La Sierra de Guadarrama, Spain. Ekologia (CSFR) 10(1): 19-29.

Papenfuss, T.J. 1960. Toad poisoning in a lizard. Herpetologica 16: 43.

Peek, R. 2017. Identification and characterization of a small sized morph of ocellated lizard (Timon lepidus Daudin, 1802) from central Spain. L@certidae 2017(1): 1-12.

Peters, G. 1961. Die Perleidechse (Lacerta lepida Daudin) gehört zum Subgenus Gallotia Boulenger. Mitteilungen aus dem Zoologischen Museum in Berlin 37: 271-285.

Peters, G. 1962. Ein Beitrag zur Ökologie der Perleidechse (Lacerta l. lepida Daudin). Mitteilungen aus dem Zoologischen Museum in Berlin 38: 401-413.

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Timon princeps

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