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Nieto-Montes de Oca, A.; Arenas-Moreno, D.M.; Beltrán-Sánchez, E.; Leaché, A.D. 2014 A new species of horned lizard (genus Phrynosoma) from Guerrero, Mexico, with an updated multilocus phylogeny. Herpetologica 70(2): 241-257.