Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk). Latest update:
October 9th, 2024.
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Genus
Calotes
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South Asian Forest Agamas, Forest Lizards, Garden Lizards, Tree Agamas, Variable Lizards, Varied Lizards, Blood-suckers
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Schönechsen
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Pragtagamer
|
1817 | |
Calotes Cuvier (type species: Lacerta calotes Linnaeus 1758) |
1856 | |
Brachysaura Blyth (type species: Brachysaura ornata Blyth 1856; syn. Deepak, Vyas, Giri & Karanth 2015) |
1864 | |
Oriocalotes Günther (type species: Calotes minor Gray 1845; syn. Giri, Chaitanya, Mahony, Lalronunga, Lalrinchhana, Das, Sarkar, Karanth & Deepak 2019) |
1870 | |
Oreocalotes Jerdon [substitute name for Oriocalotes Günther 1864] (syn. Wermuth 1965) |
1997 | |
Complicitus Manthey & Grossmann (type species: Calotes nigrigularis Ota & Hikida 1991) |
Contents:
34 species, of which 21 (61.8%) are endemic.
Endemism: 0% 100%
Remarks:
Previously included Pseudocalotes (e.g. Wermuth 1967),
and the species now assigned to
Microauris
and
Monilesaurus.
Includes C. minor, formerly assigned to a separate, monotypic genus, Brachysaura,
and C. paulus, formerly assigned to a separate, monotypic genus, Oriocalotes.
Distribution:
S. Asia, Malay Archipelago.
Reported from:
Afghanistan,
Bangladesh,
Bhutan,
Cambodia,
China
(Guangdong,
Guangxi,
Hainan,
Hong Kong,
Hunan,
Macau,
Tibet,
Yunnan),
India
(Andaman & Nicobar Islands
[Andaman Islands],
Nicobar Islands,
Andhra Pradesh,
Arunachal Pradesh,
Assam,
Bihar,
Chhattisgarh,
Delhi,
Goa,
Gujarat,
Haryana,
Himachal Pradesh,
Jammu & Kashmir,
Jharkhand,
Karnataka,
Kerala,
Lakshadweep,
Madhya Pradesh,
Maharashtra,
Manipur,
Meghalaya,
Mizoram,
Nagaland,
Odisha,
Punjab,
Rajasthan,
Sikkim,
Tamil Nadu,
Telangana,
Tripura,
Uttar Pradesh,
Uttarakhand,
West Bengal),
Indonesia
(Ambon,
Natuna Islands),
Iran,
Laos,
Malaysia
(Sabah,
West Malaysia
[incl.
Langkawi Archipelago,
Penang Island]),
Myanmar,
Nepal,
Pakistan,
Sri Lanka,
Thailand,
Vietnam
(incl.
Con Dao Islands
[incl.
Con Son]).
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Calotes bachae
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Eastern Blue Crested Agama
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Östliche Blaue Schönechse
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Østlig Blå Pragtagam
|
2013 | |
Calotes bachae Hartmann, Geissler, Poyarkov, Ihlow, Galoyan, Rödder & Böhme |
Remarks:
Previously regarded as eastern populations of mystaceus.
Distribution:
Cambodia,
Laos,
Vietnam.
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Calotes bhutanensis
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Bhutanese Tree Agama, Bhutanese Calotes, Bhutan Beaty Lizard
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Bhutan-Schönechse
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Bhutanesisk Pragtagam
|
1976 | |
Calotes bhutanensis Biswas |
Remarks:
Although reported from India by Tikader & Sharma (1992), there are no verified records from the country (Venugopal 2010).
Distribution:
Bhutan.
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Calotes calotes
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Southern Green Tree Agama, Common Green Tree Agama, Green Forest Lizard, Common Tree Agama, Southern Green Tree Agama, Sawback Agama, (Malabar Green Calotes)
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Sägerückenagame
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Sydlig Grøn Pragtagam
|
1758 | |
Lacerta calotes Linnaeus |
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Iguana calotes Laurenti 1768 |
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Agama calotes Daudin 1802 |
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Calotes calotes Wagler 1830 |
1820 | |
Agama lineata Kuhl (Wermuth 1967) |
1820 | |
Agama ophiomachus Merrem (Wermuth 1967) |
Remarks:
Reports from the Nicobar Islands are considered erroneous (Das 1999).
Presence in Gujarat (e.g., Vyas 2011) is also considered erroneous, since it was not listed for the state by Patel & Vyas (2019).
Distribution:
India
(Karnataka,
Kerala,
Lakshadweep,
Odisha,
Tamil Nadu),
Sri Lanka.
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Calotes calotes
© Rune Midtgaard
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Calotes ceylonensis
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Painted-lipped Tree Agama, Painted-lipped Lizard, Ceylon Bloodsucker
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Ceylon-Schönechse
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Rødlæbet Pragtagam
|
1887 | |
Calotes mystaceus var. ceylonensis Müller |
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Calotes ceylonensis Boulenger 1890 |
1887 | |
Calotes kelaartii Nevill (Wermuth 1967) |
1896 | |
Calotes saleoides Werner (Wermuth 1967) |
Distribution:
Sri Lanka.
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Calotes chincollium
|
Collared Forest Agama, Chin Hills Garden Lizard
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|
Chin Berge-Schönechse
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Chin Hills-pragtagam
|
2003 | |
Calotes chincollium Vindum, Win, Thin, Lwin, Shein & Tun |
Distribution:
Myanmar.
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Calotes desilvai
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Sri Lankan Spotted Tree Agama, Maculate Lizard, (De Silva's Tree Agama)
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De Silvas Schönechse
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Srilankansk Plettet Pragtagam
|
2005 | |
Calotes desilvai Bahir & Maduwage |
Distribution:
Sri Lanka.
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Calotes emma
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Forest Crested Agama, Forest Crested Lizard, Forest Garden Lizard, Black-banded Garden Lizard, Spiny-headed Forest Lizard, (Emma Forest Lizard, Light Olive Assam Garden Lizard)
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Wald-Schönechse, (Emmas Schönechse)
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Skov-pragtagam
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1845 | |
Calotes emma Gray |
1927 | |
Calotes alticristatus Schmidt (Wermuth 1967) |
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Calotes emma alticristata Taylor 1963 |
Other common names:
alticristatus: Northern Forest Crested Lizard
emma: Southern Forest Crested Lizard
Distribution:
Bangladesh,
Cambodia,
China
(Guangdong,
Guangxi,
Yunnan),
India
(Assam,
Madhya Pradesh,
Manipur,
Meghalaya,
Mizoram,
Nagaland,
Tripura),
Laos,
Malaysia
(West Malaysia
[incl.
Langkawi Archipelago]),
Myanmar,
Thailand,
Vietnam
(incl.
Con Dao Islands:
Con Son).
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Calotes emma
© Henrik Bringsøe
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Calotes farooqi
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Pakistani Crested Agama
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Pakistanische Schönechse
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Pakistansk Pragtagam
|
1993 | |
Calotes versicolor nigrigularis Auffenberg & Rehman [not Calotes nigrigularis Ota & Hikida 1991] |
1995 | |
Calotes versicolor farooqi Auffenberg & Rehman [substitute name for Calotes versicolor nigrigularis Auffenberg & Rehman 1993] |
| |
Calotes farooqi Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker 2021 |
Distribution:
Afghanistan,
Pakistan.
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Calotes geissleri
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Sagaing Blue Crested Agama
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Westburmesische Blaue Schönechse
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Vestburmesisk Blå Pragtagam
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2021 | |
Calotes geissleri Wagner, Ihlow, Hartmann, Flecks, Schmitz & Böhme |
Remarks:
Previously regarded as populations of mystaceus (Wagner, Ihlow, Hartmann, Flecks, Schmitz & Böhme 2021).
Records of mystaceus from Arunachal Pradesh, Manipur, and Mizoram are tentatively assigned to geissleri, pending confirmation.
Distribution:
India
(Arunachal Pradesh,
Manipur,
Mizoram,
Nagaland),
Myanmar,
Thailand.
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Calotes geissleri
© Henrik Bringsøe
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Calotes goetzi
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Cambodian Blue Crested Agama
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Kambodschanische Blaue Schönechse
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Cambodiansk Blå Pragtagam
|
2021 | |
Calotes goetzi Wagner, Ihlow, Hartmann, Flecks, Schmitz & Böhme |
Remarks:
Previously regarded as populations of mystaceus (Wagner, Ihlow, Hartmann, Flecks, Schmitz & Böhme 2021).
Distribution:
Cambodia,
China
(Yunnan),
Laos,
Myanmar,
Thailand.
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Calotes goetzi
© Rune Midtgaard
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Calotes grandisquamis
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Large-scaled Forest Agama, Bramagherry Green Calotes
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Grossschuppen-Schönechse
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Storskællet Pragtagam
|
1875 | |
Calotes grandisquamis Günther |
Distribution:
India
(Karnataka,
Kerala,
Tamil Nadu).
|
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Calotes htunwini
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Chatthin Forest Agama, (Htunwin's Forest Lizard)
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Htun Wins Schönechse
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Chatthin-pragtagam
|
2006 | |
Calotes htunwini Zug & Vindum in Zug, Brown, Schulte & Vindum |
Distribution:
Myanmar.
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Calotes iadina
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Emerald Tree Agama
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Smaragd-Schönechse
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Smaragd-pragtagam
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2024 | |
Calotes iadina Wang, Deepak & Grismer in Wang, Deepak, Das, Grismer, Liu & Che |
Remarks:
Previously regarded as a population of jerdoni.
Distribution:
Myanmar.
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Calotes irawadi
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Ayeyarwady Forest Agama
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Ayeyarwady-Schönechse
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Ayeyarwady-pragtagam
|
2006 | |
Calotes irawadi Zug, Brown, Schulte & Vindum |
Distribution:
Bhutan,
Cambodia,
China
(Hunan,
Yunnan),
India
(Arunachal Pradesh,
Assam,
Tripura),
Myanmar,
Vietnam.
|
|
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Calotes jerdoni
|
Eastern Green Tree Agama, Montane Green Tree Agama, (Jerdon's Forest Lizard, Jerdon's Tree Agama, Khasi Hill Garden Lizard)
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Jerdons Schönechse
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Østlig Grøn Pragtagam
|
1870 | |
Calotes jerdoni Günther |
1905 | |
Calotes yunnanensis Annandale (Wermuth 1967) |
Remarks:
Records from Tibet (e.g., Zhao & Adler 1993; Ananjeva, Guo & Wang 2011) refer to
Japalura tricarinata (Schleich & Kästle (eds.) 2002; Manthey 2008).
There is a questionable, unvouchered record from Uttar Pradesh (Kanaujia, Kumar & Kumar 2017), which is disregarded herein, since all other records of this species are far to
the east of this state.
Distribution:
Bangladesh,
Bhutan,
China
(Yunnan),
India
(Arunachal Pradesh,
Assam,
Meghalaya,
Mizoram,
Nagaland),
Myanmar.
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Calotes liocephalus
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Crestless Tree Agama, Crestless Lizard, (Lionhead Agama)
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Löwenkopf-Schönechse
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Glathovedet Pragtagam
|
1872 | |
Calotes liocephalus Günther |
Remarks:
A record from Lakshadweep (Rao 1991), India, is regarded as questionable.
If the record is valid, it is most like an introduction.
Distribution:
Sri Lanka.
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Calotes liolepis
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Whistling Tree Agama, Whistling Lizard
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Sri Lanka-Schönechse
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Glatskællet Pragtagam
|
1885 | |
Calotes liolepis Boulenger |
Distribution:
Sri Lanka.
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Calotes liolepis
© Rune Midtgaard
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Calotes manamendrai
|
Riverstone Tree Agama, (Knuckles Tree Agama, Manamendra-Arachchi's Whistling Lizard)
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Riverstone-Schönechse
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Riverstone-pragtagam
|
2014 | |
Calotes manamendrai Amarasinghe, Karunarathna & Fujinuma |
Distribution:
Sri Lanka.
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Calotes maria
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Assam Forest Agama, Assam Garden Lizard, Khasi Hills Forest Lizard, (Maria's Forest Lizard)
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Khasi-Hills-Schönechse
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Assam-pragtagam
|
1845 | |
Calotes maria Gray |
1852 | |
Calotes platyceps Blyth (Wermuth 1967) |
Distribution:
Bhutan,
India
(Assam,
Meghalaya,
Mizoram,
Uttar Pradesh).
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Calotes medogensis
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Medog Forest Agama
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Medog-Schönechse
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Medog-pragtagam
|
1984 | |
Calotes medogensis Zhao & Li |
Distribution:
China
(Tibet).
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Calotes minor
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Short-tailed Ground Agama, Gujarat Dwarf Agama, Lesser Agama
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Kurzschwanz-Erdagame
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Korthalet Jordagam
|
1827 | |
Agama minor Hardwicke & Gray |
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Charasia minor Theobald 1876 |
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Acanthosaura minor Boulenger 1890 |
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Brachysaura minor Koul & Murphy 1979 |
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Laudakia minor Das 1996 |
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Calotes minor Deepak, Vyas, Giri & Karanth 2015 |
1856 | |
Brachysaura ornata Blyth (Wermuth 1967) |
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Charasia ornata Boulenger 1885 |
Remarks:
Although the type locality is in Bangladesh, presence of minor in that country is doubtful (Deepak, Vyas, Giri & Karanth 2015). A record from Odisha (Chakraborty & Dasgupta 2009) refer to Psammophilus cf. blanfordanus (Deepak, Vyas, Giri & Karanth 2015).
Not mentioned for Punjab by Singh, Dutta & Singh (2021).
Distribution:
India
(Gujarat,
Madhya Pradesh,
Punjab,
Rajasthan,
Uttar Pradesh),
Pakistan.
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Calotes mystaceus
|
Ayeyarwady Blue Crested Agama, (Western Blue Crested Agama, Blue Crested Lizard, Blue Forest Lizard, Blue Garden Lizard, Moustached Lizard, Olivaceous Garden Lizard, Indochinese Forest Lizard)
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Südburmesische Blaue Schönechse, (Westliche Blaue Schönechse)
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Sydburmesisk Blå Pragtagam, (Vestlig Blå Pragtagam)
|
1837 | |
Calotes mystaceus Dumeril & Bibron |
Remarks:
Now restricted to southern Myanmar.
Previously included the populations now assigned to bachae, geissleri, goetzi, and vindumbarbatus.
Reports from the Andaman and Nicobar Islands are considered erroneous, as no voucher specimens exist (Das 1999).
Introduced to:
USA
(Florida).
Distribution:
Myanmar.
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Calotes nemoricola
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Nilgiri Forest Agama, Nilgiri Garden Lizard
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Nilgiri-Schönechse
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Nilgiri-pragtagam
|
1853 | |
Calotes nemoricola Jerdon |
Distribution:
India
(Andhra Pradesh,
Karnataka,
Kerala,
Tamil Nadu).
|
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Calotes nigrigularis
|
Black-throated Forest Agama, Kinabalu Forest Agama
|
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Schwarzkehl-Taschenagam
|
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Sortstrubet Pragtagam
|
1991 | |
Calotes nigrigularis Ota & Hikida |
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Complicitus nigrigularis Manthey & Grossmann 1997 |
Remarks:
Status uncertain. Some authors have placed it in a separate, monotypic genus, Complicitus
(e.g., Manthey & Grossmann 1997; Manthey & Denzer 2000; Iskandar & McGuire 2018).
Distribution:
Malaysia
(Sabah).
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Calotes nigrilabris
|
Black-lipped Tree Agama, Black-lipped Lizard
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Schwarzlippen-Schönechse
|
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Sortlæbet Pragtagam
|
1860 | |
Calotes nigrilabris Peters |
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Otocryptis nigrilabris Peters 1864 |
Distribution:
Sri Lanka.
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Calotes nigrilabris
© Henrik Bringsøe
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Calotes nigriplicatus
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Black-shouldered Tree Agama
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Schwarzschulter-Schönechse
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Sortskuldret Pragtagam
|
2000 | |
Calotes nigriplicatus Hallerman |
Distribution:
Indonesia
(Ambon).
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|
Calotes paulus
|
Northern Lesser Forest Agama, Pygmy Forest Lizard, Small Forest Lizard, Dwarf Calotes, (Khasi Hills Forest Lizard)
|
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Nördlicher Zwergwaldagame
|
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Nordlig Dværgskovagam
|
1845 | |
Calotes minor Gray [misinterpretation of Agama minor Hardwicke & Gray 1827] (Wermuth 1967) |
| |
Acanthosaura minor Boulenger 1885 |
1935 | |
Oriocalotes paulus Smith |
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Calotes paulus Giri, Chaitanya, Mahony, Lalronunga, Lalrinchhana, Das, Sarkar, Karanth & Deepak 2019 |
Remarks:
Currently known with certainty only from Khasi Hills, Meghalaya, India (Giri, Chaitanya, Mahony, Lalronunga, Lalrinchhana, Das, Sarkar, Karanth & Deepak 2019).
Several other distribution records have been published, but they all need confirmation.
These include Tibet (Zhao & Adler 1993; Rao 2000; Ananjeva, Guo & Wang 2011), Arunachal Pradesh, India (Sanyal & Gayen 2006), Assam, India (Tikader & Sharma 1992),
and Sikkim, India (Zhao & Adler 1993).
Distribution:
India
(Meghalaya).
|
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|
Calotes pethiyagodai
|
Midland Tree Agama, (Knuckles Crestless Agama, Pethiyagoda's Crestless Lizard)
|
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Midland-Schönechse
|
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Midland-pragtagam
|
2014 | |
Calotes pethiyagodai Amarasinghe, Karunarathna, Hallermann, Fujinuma, Grillitsch & Cambell |
Distribution:
Sri Lanka.
|
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Calotes versicolor
|
South Indian Crested Agama, Oriental Crested Agama, Oriental Garden Lizard, Indian Garden Lizard, Indian Bloodsucker, Garden Fence Lizard, Changeable Lizard, Red-headed Lizard
|
|
Südindische Schönechse, Verschiedenfarbige Schönechse, Gewöhnliche Schönechse, Blutsaugeragame
|
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Sydindisk Pragtagam, Almindelig Pragtagam, Indisk Pragtagam
|
1802 | |
Agama versicolor Daudin |
| |
Calotes versicolor Fitzinger 1826 |
1820 | |
Agama tiedemanni Kuhl (Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker 2021) |
1844 | |
Calotes cristatus Jaqucmont (Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker 2021) |
1846 | |
Calotes viridis Gray (Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker 2021) |
Remarks:
Restricted to southern India by Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker (2021), which leaves the remaining populations as taxonomically unresolved.
Records of versicolor from these populations are retained as versicolor herein, pending further studies to clarify their status.
Previously included farooqi as well as the populations now assigned to vultuosus and wangi.
Known from a single specimen from Oman, which was probably introduced (Arnold 1986).
Records from Afghanistan (e.g., Wermuth 1967; Leviton & Anderson 1970; Sindaco & Jeremchenko 2008; Wagner, Bauer, Leviton, Wilms & Böhme 2016)
are provisionally referred to farooqi, pending morphological and genetic confirmation.
Records from Pakistan, outside the range of farooqi, and Iran are provisionally retained as versicolor, although these populations are likely to represent other species
(see Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker 2021).
Introduced to:
Brunei,
Chagos Archipelago,
Indonesia
(Sulawesi,
Sumatra),
Kenya,
Maldives,
Mauritius,
New Zealand,
Oman,
Reunion,
Seychelles,
Singapore,
USA
(Florida).
Distribution:
Bangladesh,
Bhutan,
Cambodia,
China
(Fujian,
Guangdong,
Guangxi,
Hainan,
Hong Kong,
Hunan,
Macau,
Yunnan),
India
(Andaman & Nicobar Islands
[Andaman Islands,
Nicobar Islands],
Andhra Pradesh,
Arunachal Pradesh,
Assam,
Bihar,
Chhattisgarh,
Delhi,
Goa,
Gujarat,
Haryana,
Himachal Pradesh,
Jammu & Kashmir,
Jharkhand,
Karnataka,
Kerala,
Lakshadweep,
Madhya Pradesh,
Maharashtra,
Manipur,
Meghalaya,
Mizoram,
Nagaland,
Odisha,
Punjab,
Rajasthan,
Sikkim,
Tamil Nadu,
Telangana,
Tripura,
Uttar Pradesh,
Uttarakhand,
West Bengal),
Indonesia
(Natuna Islands),
Iran,
Laos,
Malaysia
(West Malaysia
[incl.
Penang Island]),
Myanmar,
Nepal,
Pakistan,
Sri Lanka,
Thailand,
Vietnam.
|
Calotes versicolor
© Rune Midtgaard
|
|
Calotes vindumbarbatus
|
Kachin Blue Crested Agama
|
|
Nordburmesische Blaue Schönechse
|
|
Nordburmesisk Blå Pragtagam
|
2021 | |
Calotes vindumbarbatus Wagner, Ihlow, Hartmann, Flecks, Schmitz & Böhme |
Remarks:
Previously regarded as populations of mystaceus (Wagner, Ihlow, Hartmann, Flecks, Schmitz & Böhme 2021).
Distribution:
Myanmar.
|
|
|
Calotes vultuosus
|
Kolkata Crested Agama
|
|
Kolkata-Schönechse
|
|
Kolkata-pragtagam
|
1825 | |
Agama vultuosa Harlan |
| |
Calotes vultuosus Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker 2021 |
1827 | |
Agama indica Gray (syn. Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker 2021) |
Remarks:
Previously regarded as a synonym of versicolor (e.g., Wermuth 1967).
Revalidated as a separate species by Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker (2021).
Distribution:
India
(Andhra Pradesh,
Chhattisgarh,
Gujarat,
Jharkhand,
Karnataka,
Madhya Pradesh,
Maharashtra,
Rajasthan,
Telangana,
West Bengal).
|
|
|
Calotes wangi
|
Chinese Crested Agama, (Wang's Garden Lizard)
|
|
Chinesische Schönechse
|
|
Kinesisk Pragtagam
|
2023 | |
Calotes wangi Huang, Li, Wang, Li, Hou & Cai |
2023 | |
Calotes wangi hainanensis Huang, Li, Wang, Li, Hou & Cai |
Remarks:
Previously regarded as a population of versicolor (e.g., Smith 1935; Zhao & Adler 1993).
Distribution:
China
(Fujian,
Guangdong,
Guangxi,
Hainan,
Hong Kong,
Hunan,
Macao),
Vietnam.
|
|
|
Calotes yunnanensis
|
Yunnan Green Tree Agama
|
|
Yunnan-Schönechse
|
|
Yunnan-pragtagam
|
1905 | |
Calotes yunnanensis Annandale |
Remarks:
Previously regarded as a synonym of jerdoni (e.g., Wermuth 1967).
Revalidated by Wang, Deepak, Das, Grismer, Liu & Che (2024).
Distribution:
China
(Yunnan),
Myanmar.
|
|
|
Calotes zolaiking
|
Mizoram Lesser Forest Agama, Mizoram Montane Forest Lizard
|
|
Mizoram-Zwergwaldagame
|
|
Mizoram-dværgskovagam
|
2019 | |
Calotes zolaiking Giri, Chaitanya, Mahony, Lalrounga, Lalrinchhana, Das, Sarkar, Karanth & Deepak |
Distribution:
India
(Mizoram).
|
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|