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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update: December 31st, 2022.


Taxonomy of the family Viperidae
Bibliography of the genus Parias
Biodiversity of the family Viperidae








Genus
Parias

Malay Archipelago Pitvipers

Malaiische Bambusottern

Malayiske Træhugorme

1849 Parias Gray (type species: Megaera flavomaculata Gray 1842)
Contents: 10 species, of which 8 (80.0%) are endemic.
Endemism: 0% 100%
Remarks: Previously included in Trimeresurus (e.g., McDiarmid, Campbell & Touré 1999). Revalidated by Malhotra & Thorpe (2004).
Distribution: SE. Asia, Malay Archipelago.
Reported from: Brunei, Indonesia (Bangka, Batu Islands, Kalimantan, Nias, Siberut, Simeulue, Sipura, Sumatra), Malaysia (Sabah, Sarawak, West Malaysia), Philippines (Agutayan, Babuyan Islands [Babuyan Claro, Calayan, Camiguin Norte, Dalupiri], Balabac, Balete, Batan Islands [Batan, Sabtang], Biliran, Bohol, Camiguin Sur, Catanduanes, Dinagat, Leyte, Luzon, Mindanao, Mindoro, Negros, Palawan, Panay, Polillo, Siquijor, Sulu Islands [Jolo]), Singapore, Thailand.

Parias calamitas

Nias Pitviper

Nias-Bambusotter

Nias-træhugorm

2022 Trimeresurus calamitas Vogel, David & Sidik
Parias calamitas Midtgaard 2022 (this website)

Remarks: Previously regarded as a population of hageni.
Distribution: Indonesia (Nias).





Parias flavomaculatus

Yellow-spotted Pitviper, (Philippine Pitviper, Philippine Habu)

Philippinen-Bambusotter

Gulplettet Træhugorm

1842 Megaera flavomaculata Gray
Parias flavomaculatus Gray 1849
Trimeresurus flavomaculatus Günther 1879
Lachesis flavomaculatus Boulenger 1896
1842 Megaera ornata Gray (Golay & al. 1993)
Parias ornata Gray 1849
1842 Megaera variegata Gray (Golay & al. 1993)
Parias variegata Gray 1849
1885 Trimeresurus schadenbergi Fischer (Golay & al. 1993)
1910 Trimeresurus halieus Griffin (Gumprecht 2001, 2002)
Trimeresurus flavomaculatus halieus Leviton 1961

Other common names:
halieus: Polillo Pitviper
Remarks: Previously included mcgregori (see remarks under that species). Leviton, Siler, Weinell & Brown (2018) did not include Bohol in the distribution of the species.
Distribution: Philippines (Agutayan, Babuyan Islands [Babuyan Claro, Calayan, Camiguin Norte, Dalupiri], Balete, Biliran, Bohol, Camiguin Sur, Catanduanes, Dinagat, Leyte, Luzon, Mindanao, Mindoro, Negros, Panay, Polillo, Siquijor, Sulu Islands [Jolo]).


Parias gunaleni

Green-eyed Sumatran Pitviper, (Gunalen's Pitviper)

Grünaugen-Sumatra-Bambusotter

Grønøjet Sumatra-træhugorm

2014 Trimeresurus gunaleni Vogel, David & Sidik

Distribution: Indonesia (Sumatra).


Parias hageni

Bangka Green Pitviper, (Hagen's Pitviper, Hagen's Green Pitviper)

Bangka-Bambusotter, (Hagens Bambusotter)

Bangka-træhugorm

1886 Bothrops Hageni Lidth de Jeude
Trimeresurus hageni Brongersma 1933
Parias hageni Malhotra & Thorpe 2004

Remarks: Previously included the populations now assigned to calamitas, kirscheyi, and whitteni. This species has often been confused with sumatranus and possibly malcolmi in the literature (Iskandar & Colijn 2001; Gumprecht, Tillack, Orlov, Captain & Ryabov 2004). Records of sumatranus from the Indonesian islands of Bangka, Belitung, Nias, Simeulue, and some islands in the Mentawai Archipelago, are now referred to the hageni group (Vogel, David & Sidik 2014). Possibly also present in Sipora from where Boulenger (1894) and Dring, McCarthy & Whitten (1990) reported a pitviper under the name Trimeresurus sumatranus. Two specimens from Batu Island bears some similarities with whitteni, but their taxonomic status remain unresolved. Present in the past, but not confirmed recently from Singapore (Baker & Lim 2008). A record from Borneo (Soedjito 1996) may refer to a misidentified malcolmi (Iskandar & Colijn 2001).
Distribution: Indonesia (Bangka, Sumatra), Malaysia (West Malaysia), Singapore, Thailand.


Parias hageni
© Rune Midtgaard

Parias kirscheyi

Simeulue Pitviper

Simeulue-Bambusotter

Simeulue-træhugorm

2022 Trimeresurus kirscheyi Vogel, David & Sidik
Parias kirscheyi Midtgaard 2022 (this website)

Remarks: Previously regarded as a population of hageni.
Distribution: Indonesia (Simeulue).





Parias malcolmi

Mount Kinabalu Pitviper, (Malcolm’s Pitviper)

Kinabalu-Bambusotter, (Malcolms Bambusotter)

Kinabalu-træhugorm

1938 Trimeresurus sumatranus malcolmi Loveridge
Trimeresurus malcolmi Stuebing & Inger 1998
Parias malcolmi Malhotra & Thorpe 2004

Distribution: Indonesia (Kalimantan), Malaysia (Sabah, Sarawak).


Parias mcgregori

Batan Islands Pitviper, (McGregor’s Pitviper)

Batanes-Bambusotter

Batan-træhugorm

1919 Trimeresurus mcgregori Taylor
Trimeresurus flavomaculatus mcgregori Leviton 1961
Parias flavomaculatus mcgregori Malhotra & Thorpe 2004b
Parias mcgregori Malhotra & Thorpe 2004b

Remarks: Validity of mcgregori as a distinct species has been questioned (e.g., Vogel 2006; David 2007). It is treated herein as a valid species, following several recent publications (e.g., Gumprecht 2001; David, Vogel & Dubois 2011; Oliveiros, Ota, Crombie & Brown 2011; David & Vogel in Visser 2015). Malhotra & Thorpe (2004b) listed mcgregori both as a subspecies of flavomaculatus and as a separate species, but stated that the two forms are "likely to prove to be distinct species". A record from Calayan (Babuyan Islands) is regarded as a misidentification, pending further studies (Oliveiros, Ota, Crombie & Brown 2011). Leviton, Siler, Weinell & Brown (2018) did not include Sabtang (Batan Islands) in the distribution of the species.
Distribution: Philippines (Batan Islands [Batan, Sabtang]).


Parias schultzei

Palawan Pitviper, (Schultze's Pitviper)

Palawan-Bambusotter, (Schultzes Bambusotter)

Palawan-træhugorm

1909 Trimeresurus schultzei Griffin
Parias schultzei Malhotra & Thorpe 2004

Distribution: Philippines (Balabac, Palawan).


Parias sumatranus

Grey-eyed Sumatran Pitviper, Grey-tailed Green Pitviper, (Sumatran Pitviper)

Grauaugen-Sumatra-Bambusotter

Gråøjet Sumatra-træhugorm

1822 Coluber sumatranus Raffles
Coluber sumatrensis Boie 1826 [unjustified emendation]
Trimeresurus sumatranus Gray 1842
Trigonocephalus sumatranus Cantor 1847
Bothrops sumatranus Lidth de Jeude 1886
Lachesis sumatranus Boulenger 1896
Parias sumatranus Malhotra & Thorpe 2004
1844 Trigonocephalus formosus Schlegel in Müller & Schlegel (Golay & al. 1993)
Trimeresurus formosus Gray 1849
Bothrops formosus Jan 1863

Remarks: Previously included malcolmi and the population now known as gunaleni. This species has often been confused with hageni in the literature (Dring, McCarthy & Whitten 1990; Gumprecht, Tillack, Orlov, Captain & Ryabov 2004); the range of sumatranus in Sumatra is wider than indicated in Sanders & al. (2004) but the records from the Indonesian islands of Bangka, Belitung, Nias, Simeulue, and the Mentawai Archipelago, are now referred to the hageni group (Vogel, David & Sidik 2014). Presence in Singapore needs confirmation (Lim & Lim 1992).
Distribution: Brunei, Indonesia (Kalimantan, Sumatra), Malaysia (Sabah, Sarawak, West Malaysia), Thailand.


Parias sumatranus
© Rune Midtgaard

Parias whitteni

Siberut Pitviper

Siberut-Bambusotter

Siberut-træhugorm

2022 Trimeresurus whitteni Vogel, David & Sidik
Parias hageni Midtgaard 2022 (this website)

Remarks: Previously regarded as a population of hageni.
Remarks: Possibly also present in Sipora from where Dring & al. (1990) reported a pitviper under the name Trimeresurus sumatranus. Two specimens from Batu Island bears some similarities with whitteni, but their taxonomic status remain unresolved.
Distribution: Indonesia (Siberut).