Reptiles of Bohol [Philippines] Related checklists:  Philippines Bibliography of the reptiles of Bohol Taxonomic history of the reptiles of Bohol Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: April 14th, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Brown & Alcala 1970 Gaulke 1996 Hallermann 2005 Genus Draco (Gliding Dragons)   Draco bimaculatus Two-spotted Gliding Dragon Manthey 2008 McGuire & Alcala 2000   Draco cyanopterus Masked Gliding Dragon Alcala 1986 [as everetti]   Draco ornatus White-spotted Gliding Dragon McGuire & Alcala 2000 McGuire & Kiew 2001   Draco reticulatus Reticulated Gliding Dragon Alcala 1986 McGuire & Alcala 2000 McGuire & Kiew 2001 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus interruptus Mindoro Anglehead Agama Manthey 2010   Gonocephalus semperi White-spotted Anglehead Agama Alcala 1986 Brown & Alcala 1970 Welton, Siler, Grismer, Diesmos, Sites & Brown 2017 Genus Hydrosaurus (Sail-tailed Agamas)   Hydrosaurus pustulatus Philippine Sail-tailed Agama Gaulke 1989  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus annulatus Mindanao Banded Bent-toed Gecko Alcala 1986 Brown & Alcala 1978 Gaulke 1996 Welton, Siler, Diesmos & Brown 2009 Welton, Siler, Linkem, Diesmos & Brown 2010   Cyrtodactylus philippinicus Common Philippine Bent-toed Gecko Welton, Siler, Linkem, Diesmos & Brown 2010 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Brown & Alcala 1978 Genus Gekko (Tokays)   Gekko gecko Tokay Brown & Alcala 1978 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Brown & Alcala 1978   Hemidactylus platyurus Flat-tailed Gecko Brown & Alcala 1978 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus insularis Philippine Slender Gecko Brown & Alcala 1978 [as typus] Zug 2010 Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus herrei White-lined Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978   Lepidodactylus planicauda Broad-tailed Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978 Genus Pseudogekko (Smooth-scaled Geckos)   Pseudogekko brevipes Orange-spotted Smooth-scaled Gecko Alcala 1986 Brown & Alcala 1978 Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015   Pseudogekko pungkaypinit Southern Philippine Smooth-scaled Gecko Alcala 1986 [as compressicorpus] Brown & Alcala 1978 [as compressicorpus] Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Huron, Realubit, Cobb, Brown, Bergmann Morinaga, Diesmos, Diesmos, Brown & Siler 2016 Ross & Lazell 1990 [as compressicorpus] Siler, Welton, Davis, Watters, Davey, Diesmos, Diesmos & Brown 2014  Family Scincidae (Skinks) Genus Brachymeles (Indo-Malayan Slender Skinks)   * Brachymeles boholensis Bohol Slender Skink Alcala 1986 Brown & Alcala 1980 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012   Brachymeles orientalis Bario Dusita Slender Skink Alcala 1986 Brown & Alcala 1980 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown & Alcala 1980 Genus Eutropis (South Asian Sun Skinks)   Eutropis caraga Misamis Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Brown & Alcala 1980 [as m. multicarinata]   Eutropis indeprensa Mindoro Sun Skink David, Pauwels, Lays & Lenglet 2006   Eutropis multifasciata Many-lined Sun Skink Brown & Alcala 1980 O'Shea, Kusuma & Kaiser 2018 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Brown & Alcala 1980 Gaulke 1996 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia auriculata Bronze Striped Tree Skink Brown & Alcala 1980   Lipinia pulchella Yellow-striped Slender Tree Skink Alcala 1986 Brown & Alcala 1956 Brown & Alcala 1980   Lipinia quadrivittata Black-striped Slender Tree Skink Alcala 1986 Brown & Alcala 1956 Brown & Alcala 1980 Genus Otosaurus (Philippine Giant Forest Skink)   Otosaurus cumingii Philippine Giant Forest Skink Alcala 1986 Brown & Alcala 1980 Brown & Rico 2009 Genus Parvoscincus (Philippine Dwarf Forest Skinks)   Parvoscincus steerei Guimaras Dwarf Forest Skink Brown & Alcala 1980 Genus Pinoyscincus (Philippine Forest Skinks)   Pinoyscincus abdictus Camiguin Forest Skink Brown & Alcala 1980   Pinoyscincus jagorii Samar Forest Skink Brown & Alcala 1980 Ross & Lazell 1990   Pinoyscincus mindanensis Mindanao Forest Skink Alcala 1986 Brown & Alcala 1980 Ross & Lazell 1990 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus acutus Sharp-nosed Forest Skink Alcala 1986 Brown & Alcala 1980 Ross & Lazell 1990   Sphenomorphus fasciatus Philippine Banded Forest Skink Alcala 1986 Brown & Alcala 1980   Sphenomorphus variegatus Philippine Black-spotted Forest Skink Alcala 1986 Brown & Alcala 1980  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus samarensis Samar Water Monitor Boehme 2003 [as V. cumingi] David, Pauwels, Lays & Lenglet 2006 [as V. cumingi] Koch, Auliya, Schmitz, Kuch & Boehme 2007 [as V. cumingi] Koch, Gaulke & Boehme 2010 Lisle 1996 [as V. cumingi] Welton, Travers, Siler & Brown 2014

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Dunson & Minton 1978 Gorman, Licht & McCollum 1981  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Iskandar & Colijn 2001 Leviton 1967 Leviton, Siler, Weinell & Brown 2018 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga angulata Leyte Cat Snake Leviton, Siler, Weinell & Brown 2018   Boiga cynodon Dog-toothed Cat Snake Leviton, Siler, Weinell & Brown 2018 Genus Calamaria (Oriental Reed Snakes)   Calamaria lumbricoidea Variable Reed Snake Alcala 1986 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus erythrura Philippine Rat Snake Iskandar & Colijn 2001 Leviton 1979 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis marenae Philippine Painted Bronzeback Iskandar & Colijn 2001 [as pictus] Leviton 1968 [as pictus] Leviton, Siler, Weinell & Brown 2018 Vogel & Rooijen 2008   Dendrelaphis philippinensis Mindanao Stripe-tailed Bronzeback Iskandar & Colijn 2001 [as terrificus] Leviton 1968 [as terrificus] Leviton, Siler, Weinell & Brown 2018 Rooijen & Vogel 2012 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Leviton, Siler, Weinell & Brown 2018 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis auriculatus White-lined Keelback Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton 1970 Leviton, Siler, Weinell & Brown 2018   Rhabdophis lineatus Philippine Striped Keelback Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Zhu, Wang, Takeuchi & Zhao 2014 Genus Stegonotus (Indo-Australian Ground Snakes)   Stegonotus muelleri Philippine Ground Snake Barnes & Knierim 2018 Genus Tropidonophis (Indo-Australian Keelbacks)   Tropidonophis dendrophiops Mindanao Keelback Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Malnate & Underwood 1988  Family Cyclocoridae (Philippine Burrowing Snakes, etc.) Genus Oxyrhabdium (Philippine Burrowing Snakes)   Oxyrhabdium modestum Philippine Bandless Burrowing Snake Alcala 1986 Gaulke & Operiano 2006 Iskandar & Colijn 2001 Leviton 1957 Leviton 1964 Leviton, Siler, Weinell & Brown 2018  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Calliophis (South Asian Coral Snakes)   Calliophis philippinus Philippine Striped Coral Snake Brown, Smart, Leviton & Smith 2018 Genus Naja (Typical Cobras)   Naja samarensis Southeastern Philippine Spitting Cobra Alcala 1986 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Wuster & Thorpe 1991 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Leviton, Siler, Weinell & Brown 2018 Sy 2017  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Gorman, Licht & McCollum 1981 Leviton, Siler, Weinell & Brown 2018   Laticauda semifasciata Erabu Sea Krait Leviton, Siler, Weinell & Brown 2018  Family Gerrhopilidae (Indo-Malayan Blind Snakes) Genus Gerrhopilus (Indo-Malayan Blind Snakes)   Gerrhopilus hedraeus Negros Blind Snake Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Wallach 1996 Wynn 1988 [as Typhlops ater]  Family Homalopsidae (Australasian Mud Snakes) Genus Cerberus (Dog-faced Water Snakes)   Cerberus schneiderii South East Asian Dog-faced Water Snake Gorman, Licht & McCollum 1981 [as C. rynchops] Leviton, Siler, Weinell & Brown 2018 Murphy & Voris 2014 Murphy, Voris & Karns 2012  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018  Family Pareidae (Oriental Slug Eaters) Genus Aplopeltura (Blunt-headed Slug Eater)   Aplopeltura boa Blunt-headed Slug Eater Grismer, Diesmos, Gonzalez, Jose & Inger 2012 Leviton, Siler, Weinell & Brown 2018  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Gaulke 1996 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999  Family Typhlopidae (Typical Blind Snakes) Genus Ramphotyphlops (Western Pacific Blind Snakes)   Ramphotyphlops cumingii Cumming's Blind Snake Alcala 1986 Hahn 1980 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999  Family Viperidae (Vipers) Genus Parias (Malay Archipelago Pitvipers)   Parias flavomaculatus Yellow-spotted Pitviper Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Iskandar & Colijn 2001 Leviton 1963 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus subannulatus North Philippine Temple Pitviper Gaulke 1996 [as wagleri] Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 [as wagleri] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Vogel, David, Lutz, Rooijen & Vidal 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle David, Pauwels, Lays & Lenglet 2006 Diesmos, Brown, Alcala & Sison 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021