Reptiles of Benin    Bibliography of the reptiles of Benin Taxonomic history of the reptiles of Benin Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Cynisca (West African Round-headed Worm Lizards)   Cynisca leucura White-tailed Worm Lizard Chirio 2009 Gans 1987 Gans 2005 Loveridge 1952 Mane & Trape 2015 Ullenbruch, Grell & Boehme 2010

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus suchus West African Crocodile Chirio 2009 Fuchs, Mertens & Wermuth 1974 King & Burke 1989 [as niloticus] Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Wermuth & Mertens 1977 Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus tetraspis West African Dwarf Crocodile King & Burke 1989 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama agama Common Rainbow Agama Chirio 2009 Leache, Grummer, Miller, Krishnan, Fujita, Boehme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Roedel & Wagner 2016 [as A. picticauda] Loveridge 1952 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Agama gracilimembris Lesser Ground Agama Chirio & LeBreton 2007 Grandison 1968 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Wagner & Bauer 2011 Wermuth 1967   Agama parafricana Lesser African Forest Agama Leache, Grummer, Miller, Krishnan, Fujita, Boehme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Roedel & Wagner 2016 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo africanus Sahel Chameleon Ullenbruch, Grell & Boehme 2010   Chamaeleo gracilis Graceful Chameleon Chirio 2009 Loveridge 1952 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Chamaeleo necasi Togo Chameleon Glaw 2015 Tilbury 2010 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Ullenbruch, Krause & Boehme 2007   Chamaeleo senegalensis Senegal Chameleon Chirio 2009 Glaw 2015 Grandison 1956 Loveridge 1952 Tilbury 2010 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010  Family Eublepharidae (Eyelid Geckos) Genus Hemitheconyx (African Fat-tailed Geckos)   Hemitheconyx caudicinctus West African Fat-tailed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Chirio 2009 Loveridge 1947 Seufer, Kaverkin & Kirschner (eds.) 2005 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus albituberculatus White-tubercled Half-toed Gecko Trape, Trape & Chirio 2012   Hemidactylus albivertebralis White-lined Half-toed Gecko Trape, Trape & Chirio 2012   Hemidactylus angulatus West African House Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Chirio 2009 Loveridge 1952 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 [as angulatus/guineensis complex]   Hemidactylus ansorgii Slender Forest Gecko Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   * Hemidactylus beninensis Benin Half-toed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   * Hemidactylus lamaensis Lama Forest Half-toed Gecko Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Hemidactylus mabouia Cosmopolitan House Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Hemidactylus muriceus Prickly Half-toed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus conraui Cameroon Dwarf Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Lygodactylus fischeri Sierra Leone Dwarf Gecko Trape, Trape & Chirio 2012   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Trape, Trape & Chirio 2012  Family Gerrhosauridae (Plated Lizards) Genus Broadleysaurus (Rough-scaled Plated Lizard)   Broadleysaurus major Rough-scaled Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Broadley 1987 Loveridge 1952 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus boueti Benin Fringe-toed Lizard Arnold 1983 Loveridge 1952 Salvador 1982 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Genus Heliobolus (Bushveld Lizards)   Heliobolus nitida Red-tailed Bushveld Lizard Chirio 2009 Loveridge 1952 Szczerbak 1975 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Genus Holaspis (Blue-tailed Gliding Lizards)   Holaspis guentheri Western Blue-tailed Gliding Lizard Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010  Family Phyllodactylidae (Leaf-toed Geckos) Genus Ptyodactylus (Fan-toed Geckos)   Ptyodactylus togoensis Sahelian Fan-toed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 [as P. ragazzii] Chirio 2009 [as P. ragazzii] Metallinou, Cervenka, Crochet, Kratochvil, Wilms, Geniez, Shobrak, Brito & Carranza 2015 Trape, Trape & Chirio 2012 [as P. ragazzii] Ullenbruch, Grell & Boehme 2010 [as P. ragazzii] Genus Tarentola (Wall Geckos)   Tarentola ephippiata African Wall Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Chirio 2009 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides thierryi Togo Cylindrical Skink Chirio 2009 Greenbaum 2005 Loveridge 1952 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Genus Mochlus (African Supple Skinks)   Mochlus guineensis Guinean Supple Skink Chirio & LeBreton 2007 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Genus Panaspis (African Snake-eyed Skinks)   Panaspis togoensis Togo Snake-eyed Skink Chirio 2009 Fuhn 1972 Loveridge 1952 Medina, Bauer, Branch, Schmitz, Conradie, Nagy, Hibbitts, Ernst, Portik, Nielsen, Colston, Kusamba, Behangana, Roedel & Greenbaum 2016 Perret 1973 Schmitz, Ineich & Chirio 2005 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Chirio 2009 Loveridge 1952 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Trachylepis maculilabris Common Speckled-lipped Skink Loveridge 1952 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Trachylepis perrotetii Orange-flanked Skink Chirio 2009 Hoogmoed 1974 Loveridge 1952 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Trachylepis quinquetaeniata African Five-lined Skink Chirio 2009 Hoogmoed 1974 Koppetsch 2020 Loveridge 1952 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Bennett in Pianka, King & King (eds.) 2004 Chirio & LeBreton 2007 Chirio 2009 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Varanus niloticus Nile Monitor Bayless 2002 Boehme & Ziegler 1997 Chirio 2009 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Varanus stellatus African Forest Monitor Trape, Trape & Chirio 2012 [as V. ornatus] Ullenbruch, Grell & Boehme 2010 [as V. ornatus]

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Calabaria (Calabar Boa)   Calabaria reinhardtii Calabar Boa Chirio & LeBreton 2007 Trape & Balde 2014 Genus Eryx (Sand Boas)   Eryx muelleri West African Sand Boa Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010  Family Colubridae (Typical Snakes or Colubrids) Genus Afronatrix (West African Brown Water Snake)   Afronatrix anoscopus West African Brown Water Snake Chirio & Ineich 2009 Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Wallach, Williams & Boundy 2014 Genus Bamanophis (West African Whip Snake)   Bamanophis dorri West African Whip Snake Chippaux & Jackson 2019 Hughes 2013 Schatti & Trape 2008 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hippocrepis West African Herald Snake Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 Rasmussen, Chirio & Ineich 2000 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Crotaphopeltis hotamboeia Common Herald Snake Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Dasypeltis (African Egg Eaters)   Dasypeltis confusa Diamondback Egg Eater Trape & Balde 2014 Trape & Mane 2006 Trape, Mediannikov & Trape 2012   Dasypeltis gansi Savanna Egg Eater Bates, Trape & Spawls 2013 Chirio 2009 Hughes 2013 Trape & Balde 2014 Trape & Mane 2006 Trape, Mediannikov & Trape 2012 Ullenbruch, Grell & Boehme 2010 Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) David & Ineich 1999 [as D. typus] Hughes 2013 [as D. typus] Spawls & Branch 1995 [as D. typus] Trape & Balde 2014 [as D. typus] Ullenbruch, Grell & Boehme 2010 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Hapsidophrys (Emerald Snakes)   Hapsidophrys lineatus Black-lined Green Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014   Hapsidophrys smaragdinus Common Emerald Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Meizodon regularis East African Crowned Smooth Snake Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Natriciteres variegata Variable Marsh Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Philothamnus (African Green Snakes)   Philothamnus heterodermus Emerald Green Snake Chirio & LeBreton 2007 Hughes 2013 Trape, Hughes, Collet & Mediannikov 2021   Philothamnus heterolepidotus Slender Green Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014   Philothamnus irregularis Irregular Green Snake Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Philothamnus nitidus Cameroon Green Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014   Philothamnus pobeguini Guinean Green Snake Trape & Balde 2014   Philothamnus semivariegatus Variegated Bush Snake Chirio & LeBreton 2007 Chirio 2009 [as "cf. semivariegatus"] Hughes 2013 Ullenbruch, Grell & Boehme 2010   Philothamnus smithii Smith's Bush Snake Trape & Balde 2014 Trape, Hughes, Collet & Mediannikov 2021 Genus Scaphiophis (African Hook-nosed Snakes)   Scaphiophis albopunctatus Common African Hook-nosed Snake Broadley 1994 Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus variegatus Variable Cat Snake Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Thelotornis (African Vine Snakes)   Thelotornis kirtlandii Northern Vine Snake Chirio & LeBreton 2007 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas blandingii Western Black-and-Yellow Tree Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Toxicodryas pulverulenta Western Powdered Cat Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis jamesonii Central African Green Mamba Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014   Dendroaspis viridis Western Green Mamba David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes & Barry 1969 Hughes 2013 Spawls & Branch 1995 Trape & Balde 2014 Trape & Mane 2006 Ullenbruch, Grell & Boehme 2010 Genus Elapsoidea (African Garter Snakes)   Elapsoidea semiannulata Half-banded Garter Snake Broadley 1971 Chirio 2009 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hughes 2013 Spawls & Branch 1995 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Naja (Typical Cobras)   Naja katiensis Brown Spitting Cobra David & Ineich 1999 Spawls & Branch 1995 Trape & Balde 2014   Naja melanoleuca Central African Forest Cobra Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja nigricollis Black-necked Spitting Cobra Chippaux & Jackson 2019 Chirio 2009 Hughes 2013 Spawls & Branch 1995 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Wallach, Williams & Boundy 2014   Naja savannula West African Banded Cobra Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja senegalensis Senegalese Cobra Chirio 2009 Hughes 2013 Trape & Balde 2014 Trape, Chirio, Broadley & Wuster 2009 Ullenbruch, Grell & Boehme 2010 Genus Pseudohaje (Tree Cobras)   Pseudohaje nigra Black Tree Cobra David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus modestus Western Forest Centipede Eater Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake David & Ineich 1999 Hughes 2013 Rasmussen 2005 Spawls & Branch 1995 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Wallach, Williams & Boundy 2014   Atractaspis dahomeyensis Dahomey Stiletto Snake Chippaux & Jackson 2019 Chirio 2009 David & Ineich 1999 Hughes 2013 Laurent 1950 Spawls & Branch 1995 Trape & Balde 2014 Trape & Mane 2006 Ullenbruch, Grell & Boehme 2010 Wallach, Williams & Boundy 2014   Atractaspis watsoni Sokoto Stiletto Snake Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Chirio 2009 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Boaedon lineatus Lined House Snake Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Boaedon virgatus West African House Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Genus Bothrophthalmus (Red-and-Black Striped Snakes)   Bothrophthalmus lineatus Red-and-Black Striped Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Genus Chamaelycus (African Banded Snakes)   Chamaelycus fasciatus Ituri Banded Snake Chirio & LeBreton 2007 Trape & Balde 2014 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Chirio 2009 Hughes 2004 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Dromophis praeornatus Ornate Olympic Snake Chirio 2009 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Gonionotophis (Lesser File Snakes)   Gonionotophis grantii Savanna Dwarf File Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis nototaenia Southeastern Bark Snake Broadley & Hughes 2000 Hughes 2013 Trape & Balde 2014 Genus Hormonotus (Yellow Forest Snake)   Hormonotus modestus Yellow Forest Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Limaformosa (Venus File Snakes)   Limaformosa crossi Savanna File Snake Chirio 2009 Hughes 2013 Loveridge 1939 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Villiers 1965 Villiers 1975   Limaformosa guirali White-speckled Forest File Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Genus Lycophidion (African Wolf Snakes)   Lycophidion irroratum Western Forest Wolf Snake Hughes 2013 Trape & Balde 2014   Lycophidion semicinctum Semi-annulated Wolf Snake Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 Trape & Balde 2014 Trape & Mane 2006 Ullenbruch, Grell & Boehme 2010 Genus Mehelya (Greater File Snakes)   Mehelya poensis Western Forest File Snake Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum 2018 Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014   Mehelya stenophthalmus Small-eyed File Snake Chirio & LeBreton 2007 Hughes 2013 Trape & Balde 2014 Genus Polemon (African Snake Eaters)   Polemon neuwiedi West African Snake Eater Doucet 1963 Hughes & Barry 1969 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Villiers 1975 Genus Prosymna (Shovel-snout Snakes)   Prosymna collaris Collared Shovel-snout Chirio 2009 Chirio, Ineich, Schmitz & Trape 2011 Hughes 2013 Trape & Balde 2014 [as P. greigerti]   Prosymna meleagris Speckled Shovel-snout Broadley 1980 Chirio, Ineich, Schmitz & Trape 2011 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis afroccidentalis West African Sand Snake Chirio & LeBreton 2007 [as P. sibilans] Chirio 2009 [as rukwae] Hughes 2013 [as P. sibilans] Trape & Balde 2014 [as P. sibilans] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis elegans Elegant Sand Snake Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Psammophis phillipsii West African Olive Grass Snake Hughes 2013 Trape & Balde 2014 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Ullenbruch, Grell & Boehme 2010 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Chirio & Ineich 1991 Chirio 2009 Hughes 2013 Joger 1981 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis adleri Chadian Thread Snake Chirio 2009 Hughes 2013 Trape & Balde 2014 Genus Tricheilostoma (West African Forest Thread Snakes)   Tricheilostoma bicolor Bicoloured Thread Snake Chirio 2009 Doucet 1963 [as brevicauda] Hahn & Wallach 1998 Hahn 1980 [as Leptotyphlops brevicaudus] Hughes 2013 McDiarmid, Campbell & Toure 1999 [as brevicaudus] Papenfuss 1969 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 [as Leptotyphlops brevicauda] Villiers 1965  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Chirio & LeBreton 2007 Chirio 2009 Hughes 2013 McDiarmid, Campbell & Toure 1999 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Python sebae Northern Rock Python Broadley 1984 Chippaux & Jackson 2019 Chirio 2009 Hughes 2013 Spawls & Branch 1995 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops punctatus Spotted Blind Snake Hahn 1980 Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Wallach, Williams & Boundy 2014  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Chirio 2009 David & Ineich 1999 Hughes 2013 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Bitis gabonica Gaboon Adder Chippaux & Jackson 2019 Hughes 2013 Trape & Balde 2014   Bitis nasicornis African Nose-horned Viper Hughes 2013 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010   Bitis rhinoceros Rhinoceros Viper Rasmussen 1995 Genus Causus (Night Adders)   Causus maculatus West African Night Adder Chirio 2009 David & Ineich 1999 Hughes 2013 Spawls & Branch 1995 Trape & Balde 2014 Ullenbruch, Grell & Boehme 2010 Genus Echis (Carpet Vipers)   Echis ocellatus West African Carpet Viper Chirio 2009 David & Ineich 1999 Hughes 1976 Hughes 2013 McDiarmid, Campbell & Toure 1999 Pook, Joger, Stumpel & Wuster 2009 Spawls & Branch 1995 Trape & Balde 2014 Trape 2018 Ullenbruch, Grell & Boehme 2010

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Dossa, Sinsin & Mensah 2007 King & Burke 1989 Mettler, Clyde-Brockway, Honarvar & Paladino 2019 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Dossa, Sinsin & Mensah 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Dossa, Sinsin & Mensah 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Dossa, Sinsin & Mensah 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa olivacea Sahelian Helmeted Turtle Boycott & Bourquin 2008 [as subrufa] Chirio 2009 [as subrufa] Loveridge 1952 Maran 2009 Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 [as subrufa]   Pelomedusa variabilis West African Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios adansonii Sahelian Mud Turtle Chirio 2009 Trape, Trape & Chirio 2012   Pelusios castaneus Chestnut Mud Turtle Bour, Luiselli, Petrozzi, Segniabeto & Chirio 2016 Chirio & LeBreton 2007 Chirio 2009 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010   Pelusios niger West African Black Mud Turtle Chirio & LeBreton 2007 Luiselli, Bour, Petrozzi, Akani & Segniagbeto 2018 Maran 2009 McCord, Joseph-Ouni & Bour 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012  Family Testudinidae (Tortoises) Genus Centrochelys (African Spurred Tortoise)   Centrochelys sulcata African Spurred Tortoise Petrozzi, Hema, Demaya, Benansio, Eniang, Diagne, Segniagbeto & Luiselli 2020 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Kinixys (Hinge-back Tortoises)   Kinixys erosa Forest Hinge-back Tortoise Chirio & LeBreton 2007 Luiselli & Diagne 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Kinixys homeana West African Hinge-back Tortoise Chirio & LeBreton 2007 Luiselli & Diagne 2013 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012   Kinixys nogueyi Western Savanna Hinge-back Tortoise Chirio & LeBreton 2007 Chirio 2009 King & Burke 1989 [as belliana] Loveridge 1952 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 [as belliana] Ullenbruch, Grell & Boehme 2010  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis elegans Nubian Flapshell Turtle Baker, Diagne & Luiselli 2015 Chirio & LeBreton 2007 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2017 Ullenbruch, Grell & Boehme 2010   Cyclanorbis senegalensis Senegal Flapshell Turtle Chirio & LeBreton 2007 Chirio 2009 King & Burke 1989 McCord & Joseph-Ouni 2003 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Ullenbruch, Grell & Boehme 2010 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Bonin, Devaux & Dupre 2006 King & Burke 1989 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012