Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update:
August 22nd, 2026.
 
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Genus
Bronchocela
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Long-tailed Agamas, Slender Agamas
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Langschwanzagamen
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Langhaleagamer
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1827 |  |
Bronchocela Kaup (type species: Agama cristatella Kuhl 1820) |
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1841 | |
Bronchoceles Troschel [substitute name for Bronchocela Kaup 1827] |
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1843 | |
Lophodeira Fitzinger (type species: Agama cristatella Kuhl 1820; syn. Wermuth 1967) |
Contents:
21 species, of which 15 (71.4%) are endemic.
Endemism: 0%  100%
Remarks:
Previously included in the genus
Calotes (e.g., Wermuth 1967).
Revalidated by Moody (1981) and validity confirmed by Böhme (1988).
Distribution:
SE. Asia, Malay Archipelago.
Reported from:
Brunei,
Cambodia,
India
(Nicobar Islands),
Indonesia
(Ambon,
Anambas Islands,
Bacan,
Bangka,
Banyak Islands,
Bawean Islands,
Buru,
Buton,
Gorong Archipelago,
Halmahera,
Java,
Kai Islands,
Kalimantan,
Kangean Islands,
Lingga Islands,
Madura,
Misool,
Morotai,
Natuna Islands,
[incl.
Bunguran],
Nias,
Pagai Islands
[North Pagai],
Riau Islands,
Sebuku,
Selayar,
Seram,
Simeulue,
Sipura,
Sula Islands
[Sanana],
Sulawesi,
Sumatra,
Talaud Islands,
Tanimbar Islands,
Ternate,
Weh,
Western New Guinea),
Malaysia
(Sabah,
Sarawak,
West Malaysia
[incl.
Langkawi Archipelago,
Penang Island,
Perhentian Islands,
Seribuat Archipelago
(incl.
Aur,
Pemanggil,
Tioman)]),
Myanmar,
Philippines
(Bohol,
Calamian Islands
[Busuanga,
Calauit],
Camiguin Sur,
Carabao,
Catanduanes,
Cebu,
Dinagat,
Dumaran,
Leyte,
Luzon,
Mapun,
Masbate,
Mindanao,
Mindoro,
Negros,
Palawan,
Panaon,
Panay,
Polillo,
Rapu-Rapu,
Samar,
Siargao,
Sibuyan,
Sulu Islands
[Sangasanga,
Siasi,
Sibutu,
Tumindao],
Tablas),
Singapore,
Thailand
(incl.
Koh Kut Island),
Vietnam.
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Bronchocela burmana
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Burmese Green Crested Lizard
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Burmesische Langschwanzagame
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Burmesisk Langhaleagam
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1878 | |
Bronchocela burmana Blanford |
Remarks:
Previously regarded as a synonym of cristatella (e.g., Wermuth 1965; Diong & Lim 1998).
Revalidated as a separate species by Zug, Mulcahy & Vindum (2017).
Although not yet confirmed, Thai records of cristatella from north of the Isthmus of Kra are believed to refer to burmana (Zug, Mulcahy & Vindum 2017).
Distribution:
Myanmar,
Thailand
(incl.
Koh Kut Island).
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Bronchocela burmana
© Henrik Bringsøe
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Bronchocela carestiai
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Mindoro Long-tailed Agama, Mindoro Green-crested Lizard
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Mindoro-Langschwanzagame
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Mindoro-langhaleagam
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2026 | |
Bronchocela carestiai Supsup, Pitogo, Sanguila, Hallermann, Denzer, Wood & Brown |
Remarks:
Previously regarded as a population of cristatella or marmorata (Supsup, Pitogo, Sanguila, Hallermann, Denzer, Wood & Brown 2026).
Distribution:
Philippines
(Mindoro).
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Bronchocela celebensis
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Sulawesi Long-tailed Agama, Sulawesi Crested Lizard
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Sulawesi-Langschwanzagame
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Sulawesi-langhaleagam
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1845 | |
Bronchocela celebensis Gray |
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Calotes celebensis Boulenger 1885 |
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Calotes cristatellus celebensis Boettger 1901 |
Distribution:
Indonesia
(Sulawesi).
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Bronchocela cristatella
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Javan Green Crested Agama, Bornean Bloodsucker, (Malayan Green Crested Agama, Peninsular Green Crested Lizard, Red-spotted Garden Lizard, Green Tree Lizard)
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Javanische Grüne Langschwanzagame, (Borneo-Langschwanzagame, Borneo-Schönechse)
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Javansk Grøn Langhaleagam
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1820 | |
Agama cristatella Kuhl |
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Bronchocela cristatella Kaup 1827 |
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Calotes cristatella Wagler 1830 |
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Agama cristatella Gray 1831 |
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Lophodeira cristatellus Fitzinger 1834 |
Remarks:
Now restricted to W. Indonesia.
Previously included burmana, celebensis, moluccana, octospinosa, and petersidoriai, as well as the populations now assigned to
carestiai, mallarii, marmorata, philippina, and rubrigularis.
The taxonomic status of many older records of cristatella are unclear and need further studies.
These include records from Bawean Islands and Kangean Islands which provisionally are retained herein as cristatella.
Records from Morotai, Bacan, Gorong Archipelago, Talaud Islands, Ternate, Misool, Morotai, and Sula Islands are tentatively referred to moluccana.
Records from Brunei, Sabah, Sarawak, Anambas Islands, Bangka, Banyak Islands, Natuna Islands, Natuna Islands, Nias, Pagai Islands, Riau Islands, Simeulue, Sipura, and Weh are tentatively referred
to octospinosa.
Records from Sulawesi refer to celebensis or petersidoriai (Amarasinghe, Campbell, Riyanto, Hallermann, Hamidy, Andayani, Abinawanto & Supriatna 2024;
Amarasinghe, Jablonski, Poyarkov, Campbell, Ineich, Riyanto, Trofimets, Andayani, Abinawanto, Supriatna, Hamidy & Hallermann 2026).
Records from India (Nicobar Islands) refer to rubrigularis (Hallermann 2009).
Records from Myanmar and Thai records of cristatella from north of the Isthmus of Kra are believed to refer to burmana (Zug, Mulcahy & Vindum 2017).
Philippine populations of cristatella are now assigned to carestiai, mallarii, philippina, or marmorata
(Supsup, Pitogo, Sanguila, Hallermann, Denzer, Wood & Brown 2026).
Records from Kai Islands, Tanimbar Islands, and New Guinea are not assigned at species level.
Distribution:
Indonesia
(Bali,
Bawean Islands,
Java,
Kalimantan,
Kangean Islands,
Sumatra).
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Bronchocela cristatella
© Henrik Bringsøe
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Bronchocela cyanopalpebra
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Car Nicobar Long-tailed Agama
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Car-Nicobar-Langschwanzagame
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Car Nicobar-langhaleagam
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2023 | |
Bronchocela cyanopalpebra Chandramouli, Adhikari, Amarasinghe & Abinawanto |
Distribution:
India
(Andaman & Nicobar Islands:
Nicobar Islands:
Car Nicobar).
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Bronchocela danieli
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Great Nicobar Long-tailed Agama, Great Nicobar Calotes, (Daniel's Forest Lizard)
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Nicobarische Langschwanzagame
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Nicobarisk Langhaleagam
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1973 | |
Calotes danieli Tiwari & Biswas |
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Bronchocela danieli Welch, Cooke & Wright 1990 |
Distribution:
India
(Nicobar Islands:
Great Nicobar).
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Bronchocela hayeki
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Sumatran Long-tailed Agama, (Hayek's Forest Lizard)
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Hayeks Langschwanzagame
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Sumatra-langhaleagam
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1928 | |
Calotes hayeki Müller |
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Bronchocela hayeki Welch, Cooke & Wright 1990 |
Distribution:
Indonesia
(Sumatra).
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Bronchocela jubata
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Maned Long-tailed Agama, Maned Forest Lizard, Yellow-spotted Forest Lizard, Great Crested Canopy Lizard (Nicobar Green Lizard)
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Mähnen-Langschwanzagame
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Manke-langhaleagam
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1837 | |
Bronchocela jubata Dumeril & Bibron |
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Calotes jubata Duméril & Duméril 1851 |
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1842 | |
Calotes intermedius Berthold (syn. Böhme & Bischoff 1984) |
Remarks:
Reports from the Nicobar Islands are considered erroneous, as no voucher specimens exist (Das 1999).
Likewise, a record from Puduchery (formerly Pondichery) is considered doubtful and may have been based on an introduction (Hallermann 2005).
Presence in India is thus regarded as unverified.
Also records from Cambodia, Philippines, and Sulawesi are regarded as erroneous (Amarasinghe, Ineich, Riyanto, Hallermann, Andayani, Abinawanto & Supriatna 2022).
Distribution:
Indonesia
(Bali,
Java,
Kalimantan,
Lingga Islands,
Madura,
Nias,
Sebuku,
Sumatra),
Thailand.
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Bronchocela mallarii
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Palawan Long-tailed Agama, Tau't Bato Green-crested Lizard
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Palawan Langschwanzagame
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Palawan-langhaleagam
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2026 | |
Bronchocela mallarii Supsup, Pitogo, Sanguila, Hallermann, Denzer, Wood & Brown |
Remarks:
Previously regarded as a population of cristatella (Supsup, Pitogo, Sanguila, Hallermann, Denzer, Wood & Brown 2026).
Records of cristatella from Calamian Islands, Dumaran, and Mapun are tentatively referred to mallarii.
Distribution:
Philippines
(Calamian Islands
[Busuanga,
Calauit],
Dumaran,
Mapun,
Palawan).
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Bronchocela marmorata
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Marbled Long-tailed Agama, Marmorated Crested Lizard, Marmorated Bloodsucker, Philippine Calotes
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Marmorierte Langschwanzagame
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Marmor-langhaleagam
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1845 | |
Bronchocela marmorata Gray |
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Calotes marmoratus Boulenger 1885 |
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1922 | |
Calotes marmoratus sanchezi Taylor (syn. Supsup, Pitogo, Sanguila, Hallermann, Denzer, Wood & Brown 2026) |
Remarks:
Previously included philippina and
Hypsilurus spinosus.
Distribution:
Philippines
(Carabao,
Catanduanes,
Cebu,
Luzon,
Lubang,
Marinduque,
Negros,
Panay,
Polillo,
Romblon,
Sibuyan,
Tablas).
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Bronchocela moluccana
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Moluccan Green Crested Agama
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Molukkanische Grüne Langschwanzagame
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Molukkansk Grøn Langhaleagam
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1830 | |
Agama moluccana Lesson |
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Bronchocela moluccana Amarasinghe, Jablonski, Poyarkov, Campbell, Ineich, Riyanto, Trofimets, Andayani, Abinawanto, Supriatna, Hamidy & Hallermann 2026 |
Remarks:
Previously regarded as a synonym of cristatella (e.g., Diong & Lim 1998).
Revalidated as a separate species by Amarasinghe, Jablonski, Poyarkov, Campbell, Ineich, Riyanto, Trofimets, Andayani, Abinawanto, Supriatna, Hamidy & Hallermann (2026).
Some of the distribution records below are tentatively assigned to this species based on geography, however, confirmation is needed (see remarks under cristatella).
Distribution:
Indonesia
(Ambon,
Bacan,
Buru,
Gorong Archipelago,
Halmahera,
Misool,
Morotai,
Seram,
Sula Islands
[Sanana],
Talaud Islands,
Ternate).
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Bronchocela nicobarica
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Little Nicobar Long-tailed Agama
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Little-Nicobar-Langschwanzagame
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Little Nicobar-langhaleagam
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2023 | |
Bronchocela nicobarica Chandramouli, Adhikari, Amarasinghe & Abinawanto |
Distribution:
India
(Andaman & Nicobar Islands:
Nicobar Islands
[Kondul,
Little Nicobar,
Menchal,
Pilo Milo]).
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Bronchocela octospinosa
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Malayan Green Crested Agama, Peninsular Green Crested Lizard, Green Tree Lizard, Bornean Bloodsucker, Red-spotted Garden Lizard, (Indonesian Calotes)
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Malayische Grüne Langschwanzagame, (Borneo-Langschwanzagame, Borneo-Schönechse)
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Malayisk Grøn Langhaleagam
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1913 | |
Calotes octospinosus Baumann |
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Bronchocela octospinosa Amarasinghe, Jablonski, Poyarkov, Campbell, Ineich, Riyanto, Trofimets, Andayani, Abinawanto, Supriatna, Hamidy & Hallermann 2026 |
Remarks:
Previously regarded as a synonym of cristatella (e.g., Wermuth 1967; Diong & Lim 1998).
Revalidated as a separate species by Amarasinghe, Jablonski, Poyarkov, Campbell, Ineich, Riyanto, Trofimets, Andayani, Abinawanto, Supriatna, Hamidy & Hallermann (2026).
Some of the distribution records below are tentatively assigned to this species based on geography, however, confirmation is needed (see remarks under cristatella).
Distribution:
Brunei,
Indonesia
(Anambas Islands,
Bangka,
Banyak Islands,
Kalimantan,
Natuna Islands,
Nias,
Pagai Islands
[North Pagai],
Riau Islands,
Simeulue,
Sipura,
Sumatra,
Weh),
Malaysia,
(Sabah
[incl.
Banggi Islands],
Sarawak,
West Malaysia
[incl.
Bidong Island,
Langkawi Archipelago,
Penang Island,
Perhentian Islands,
Seribuat Archipelago
(incl.
Aur,
Pemanggil,
Tioman)]),
Singapore,
Thailand.
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Bronchocela orlovi
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Gia Lai Long-tailed Agama, (Orlov's Long-tailed Agama, Orlov's Forest Lizard)
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Orlovs Langschwanzagame
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Gia Lai-langhaleagam
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2004 | |
Bronchocela orlovi Hallermann |
Distribution:
Vietnam.
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Bronchocela petersidoriai
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Kendari Green Crested Agama
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Kendari-Langschwanzagame
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Kendari-langhaleagam
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1878 | |
Bronchocela intermedia Peters & Doria [not Calotes intermedius Berthold 1842] (syn. Bischoff & Böhme 1984) |
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1984 | |
Bronchocela petersidoriai Böhme & Bischoff [substitute name for Bronchocela intermedia Peters & Doria 1878] |
Remarks:
Previously regarded as a synonym of cristatella (e.g., Bischoff & Böhme 1984).
Revalidated as a separate species by Amarasinghe, Jablonski, Poyarkov, Campbell, Ineich, Riyanto, Trofimets, Andayani, Abinawanto, Supriatna, Hamidy & Hallermann (2026).
Distribution:
Indonesia
(Buton,
Sulawesi).
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Bronchocela philippina
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Mindanao Long-tailed Agama
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Mindanao Langschwanzagame
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Mindanao-langhaleagam
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1867 | |
Calotes philippinus Peters |
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Bronchocela philippina Supsup, Pitogo, Sanguila, Hallermann, Denzer, Wood & Brown 2026 |
Remarks:
Previously regarded as a synonym of marmorata (e.g., Wermuth 1967).
Revalidated as a separate species by Supsup, Pitogo, Sanguila, Hallermann, Denzer, Wood & Brown (2026).
Records of cristatella from Panaon and the Sulu Islands are tentatively referred to philippina.
Distribution:
Philippines
(Bohol,
Camiguin Sur,
Dinagat,
Leyte,
Mindanao,
Panaon,
Samar,
Siargao,
Sulu Islands
[Sangasanga,
Siasi,
Sibutu,
Tumindao]).
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Bronchocela rayaensis
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Langkawi Long-tailed Agama, Mount Raya Green Crested Lizard
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Langkawi-Langschwanzagame
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Langkawi-langhaleagam
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2015 | |
Bronchocela rayaensis Grismer, Wood, Lee, Quah, Anuar, Ngadi & Sites |
Distribution:
Malaysia
(West Malaysia:
Langkawi Archipelago),
Thailand.
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Bronchocela rayaensis
© Henrik Bringsøe
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Bronchocela rubrigularis
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Red-throated Long-tailed Agama
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Rotkehl-Langschwanzagame
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Rødstrubet Langhaleagam
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2010 | |
Bronchocela rubrigularis Hallermann |
Remarks:
Previously regarded as a population of cristatella (e.g., Harikrishnan, Vasudevan & Choudhury 2010; Das 1996, 2003; Manthey 2008; Sharma 2002; Venugopal 2010a).
Distribution:
India
(Nicobar Islands
[Bompoka,
Camorta,
Chowra,
Katchall,
Nancowry,
Teressa,
Tillanchong,
Trinket]).
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Bronchocela shenlong
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Mount Larut Long-tailed Agama, Shen Long Green Crested Lizard
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Larut-Langschwanzagame
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Larut-langhaleagam
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2015 | |
Bronchocela shenlong Grismer, Wood, Lee, Quah, Anuar, Ngadi & Sites |
Distribution:
Malaysia
(West Malaysia).
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Bronchocela smaragdina
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Emerald Long-tailed Agama, (Chantaburi Green Tree Lizard, Chantaburi Forest Lizard, Günther's Bloodsucker)
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Smaragd-Langschwanzagame
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Smaragd-langhaleagam
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1864 | |
Bronchocela smaragdina Günther |
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Calotes smaragdinus Boulenger 1885 |
Remarks:
Presence in Thailand needs confirmation (Pauwels & David 2005; Hallermann 2005), although reported from Chanthaburi by several authors (e.g., Nabhitabhata, Chan-ard & Chuaynkern 2004; Chuaynkern & Chuaynkern 2012; Chan-ard, Parr & Nabhitabhata 2015).
Distribution:
Cambodia,
Vietnam.
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Bronchocela vietnamensis
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Vietnamese Long-tailed Agama, Vietnamese Forest Lizard
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Vietnamesische Langschwanzagame
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Vietnamesisk Langhaleagam
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2005 | |
Bronchocela vietnamensis Hallermann |
Distribution:
Vietnam.
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