Home - Taxonomy - Geography - Biodiversity - Literature - Purchase RepFocus Recent updates Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk). Latest update: December 31st, 2022. Taxonomy of the family Gekkonidae Bibliography of the genus Gehyra Biodiversity of the family Gekkonidae Genus Gehyra Four-clawed Geckos, Web-toed Geckos, Dtellas Vierkrallengeckos Stumpfingergekkoer 1834 Gehyra Gray (type species: Gecko oceanicus Lesson 1830) 1835 Peropus Wiegmann (type species: Hemidactylus mutilatus Wiegmann 1835; syn. Bauer 1994) 1842 Phyria Gray (type species: Phyria punctulata Gray 1842; syn. Bauer 1994) 1843 Dactyloperus Fitzinger (type species: Hemidactylus variegatus Dumeril & Bibron 1836; syn. Bauer 1994) 1843 Perodactylus Fitzinger (type species: Hemidactylus oualensis Dumeril & Bibron 1836; syn. Bauer 1994) 1845 Peripia Gray (type species: Hemidactylus peronii Dumeril & Bibron 1836; syn. Bauer 1994) 1985 Phyriadoria Wells & Wellington (type species: Gehyra australis Gray 1845; syn. Bauer 1994) Contents: 69 species, of which 62 (89.9%) are endemic. Endemism: 0% 100% Distribution: Madagascar, S. Asia, Malay Archipelago, Australia, Indian Ocean, Pacific Ocean. Reported from: American Samoa, Australia (New South Wales, Northern Territory, Queensland [incl. Fraser Island, Torres Strait Islands], South Australia, Victoria, Western Australia), Brunei, Cambodia, China (Guangdong, Hainan, Hong Kong, Yunnan), Clipperton, Cocos (Keeling) Islands, Cook Islands (incl. Rarotonga), East Timor (incl. Atauro Island), Easter Island, Fiji (incl. Rotuma), French Polynesia, Guam, India (Andaman & Nicobar Islands [Andaman Islands, Nicobar Islands], Kerala), Indonesia (Alor, Ambon, Anambas Islands, Aru Islands, Bacan, Bali, Banda Islands, Banggai Islands, Bangka, Batanta, Batu Islands, Buru, Buton, Ende, Enggano, Flores, Gorong Archipelago, Halmahera, Hoga, Java, Kabaena, Kai Islands, Kalimantan, Kangean Islands, Karimunjawa Islands, Komodo, Krakatau, Legundi, Lombok, Morotai, Natuna Islands, Nias, Obi, Panaitan, Penida, Rote, Salawati, Sangir Islands [incl. Biaro], Saparua, Sebesi, Selayar, Sula Islands, Sulawesi, Sumatra, Sumba, Sumbawa, Supiori, Tanahjampea, Tanimbar Islands, Ternate, Tinjil, Waigeo, West Timor, Western New Guinea, Wetar), Japan (Ryukyu Islands [Okinawa Islands, Sakishima Islands, Satsunan Islands]), Kiribati, Laos, Madagascar (incl. Nosy Be), Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Penang Island, Perhentian Islands, Seribuat Archipelago (incl. Aur, Pemanggil, Tioman)]), Marshall Islands (incl. Arno Atoll), Micronesia (incl. Pohnpei [incl. Ant Atoll]), Myanmar, Nauru, New Caledonia (incl. Loyalty Islands), Niue, Northern Mariana Islands, Palau, Papua New Guinea (incl. Admiralty Islands [Manus], Bismarck Archipelago [New Britain, New Ireland, Umboi], d'Entrecasteaux Islands [Fergusson, Normanby], Eastern New Guinea, Louisiade Archipelago [Rossel, Sudest], Woodlark, North Solomon Islands [Bougainville]), Philippines (Apo, Babuyan Islands [Calayan, Camiguin Norte, Dalupiri, Fuga], Balabac, Balicasag, Bantayan, Batan Islands [Batan], Bohol, Bonoon, Boracay, Calagnaan, Calamian Islands [Calauit], Caluya, Camiguin Sur, Camotes Islands [Pacijan, Ponson, Poro], Carabao, Catanduanes, Cebu, Clara, Danjugan, Dinagat, Duitay, Dumaran, Gigante Islands, Guimaras, Inampulugan, Jao, Lapinig Islands [Lapinig Chico, Lapinig Grande], Leyte, Luzon, Mactan, Mantique, Marinduque, Mindanao, Mindoro, Nadulao, Negros, Palawan, Pan de Azucar, Panay, Panubulon, Samar, Sibuyan, Sicogon, Siquijor, Sulu Islands [Bongao, Bubuan, Jolo], Tablas, Tilmubo), Pitcairn Islands, Seychelles (incl. Aldabra, Praslin), Singapore, Solomon Islands (incl. Bellona, Taumako, Guadalcanal, Rennell, Santa Cruz Islands), Sri Lanka, Taiwan, Thailand, Tokelau, Tonga, Tuvalu, Vanuatu (incl. Espiritu Santo), Vietnam, Wake, Wallis & Futuna, Western Samoa. Gehyra angusticaudata Narrow-tailed Four-clawed Gecko, Slender-tailed Four-clawed Gecko Dünnschwanz-Vierkrallengecko Tyndhalet Stumpfingergekko 1963 Peropus angusticaudatus Taylor Gehyra angusticaudata Wermuth 1965 Distribution: Thailand. Gehyra arnhemica East Arnhem Land Dtella Östlicher Arnhemland-Vierkrallengecko Østlig Arnhemland-stumpfingergekko 2020 Gehyra arnhemica Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz Distribution: Australia (Northern Territory [incl. Groote Eylandt]). Gehyra australis Western Top End Dtella, (Northern Dtella, Australian House Gecko) Port-Essington-Vierkrallengecko, (Australischer Hausgecko) Port Essington-stumpfingergekko, (Nordaustralsk Stumpfingergekko) 1845 Gehyra australis Gray Hemidactylus australis Schmeltz 1874 Peropus variegatus australis Loveridge 1934 Gehyra variegata australis Mitchell 1955 Phyriadoria australis Wells & Wellington 1985 1842 Phyria punctulata Gray 1842 [nomen oblitum] (Bauer 1994) Remarks: Validity as a separate species confirmed by Bustard (1965). Now restricted to Northern Territory. Previously included the populations now assigned to arnhemica, gemina, and lauta. Records from Queensland and Western Australia refer to these species (Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz 2020). Distribution: Australia (Northern Territory). Gehyra australis © Rune Midtgaard Gehyra baliola Short-tailed Dtella Kurzschwanz-Vierkrallengecko Korthalet Stumpfingergekko 1851 Hemidactylus baliolus Dumeril Gehyra baliola Boulenger 1885 Peropus baliolus Chrapliwy, Smith & Grant 1961 1877 Peripia brevicaudis Macleay (Bauer 1994) 1877 Peripia marmorata Macleay (Bauer 1994) 1877 Peripia torresiana Günther (Bauer 1994) Gehyra torresiana Wells & Wellington 1985 Distribution: Australia (Queensland [incl. Torres Strait Islands]), Indonesia (Waigeo), Papua New Guinea (Eastern New Guinea). Gehyra barea Banda Islands Dtella Banda-Vierkrallengecko Banda-stumpfingergekko 1926 Gehyra barea Kopstein Peropus barea Chrapliwy, Smith & Grant 1961 Remarks: Suggested to be synonymous with membranacruralis by Bauer & Henle (1995), but validity confirmed by Oliver, Sistrom, Tjaturadi, Krey & Richards (2010). Distribution: Indonesia (Banda Islands, Batanta, Salawati). Gehyra borroloola Borroloola Dtella Borroloola-Vierkrallengecko Borroloola-stumpfingergekko 1983 Gehyra borroloola King Dactyloperus borroloola Wells & Wellington 1985 Phyriadoria borroloola Wells & Wellington 1989 Distribution: Australia (Northern Territory, Queensland). Gehyra brevipalmata Palau Dtella, (Palau Gecko) Palau-Vierkrallengecko Palau-stumpfingergekko 1874 Hemidactylus brevipalmatus Peters Gehyra brevipalmata Boulenger 1885 Peropus brevipalmatus Chrapliwy, Smith & Grant 1961 Distribution: Palau. Gehyra calcitectus Relictual Karst Dtella Karst-Vierkrallengecko Karst-stumpfingergekko 2020 Gehyra calcitectus Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz Distribution: Australia (Western Australia). Gehyra capensis North West Cape Dtella, North West Cape Gehyra North-West-Cape-Vierkrallengecko North West Cape-stumpfingergekko 2018 Gehyra capensis Kealley, Doughty, Pepper, Keogh, Hillyer & Huey Distribution: Australia (Western Australia). Gehyra catenata Chain-backed Dtella, (Tree Dtella, Tree Gecko) Ketten-Vierkrallengecko Kædemønstret Stumpfingergekko 1979 Gehyra catenata Low Dactyloperus catenata Wells & Wellington 1985 Distribution: Australia (Queensland). Gehyra chimera Western Kimberley Tree Dtella Chimären-Vierkrallengecko Kimære-stumpfingergekko 2020 Gehyra chimera Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz Distribution: Australia (Western Australia). Gehyra crypta Western Variegated Dtella, Western Cryptic Gehyra Westlicher Bunter Vierkrallengecko Vestlig Broget Stumpfingergekko 2018 Gehyra crypta Kealley, Doughty, Pepper, Keogh, Hillyer & Huey Distribution: Australia (Western Australia). Gehyra dubia Eastern Dtella, Dubious Dtella, (Northern Dtella, Australian House Gecko) Ostaustralischer Vierkrallengecko Østaustralsk Stumpfingergekko 1877 Peripia dubia Macleay Gehyra dubia King 1985 Dactyloperus dubia Wells & Wellington 1985 Phyriadoria dubia Wells & Wellington 1989 1867 Gecko grayi Steindachner (Bauer 1994) Dactyloperus grayi Wells & Wellington 1984 1877 Peripia longicaudis Macleay (Bauer 1994) Dactyloperus longicaudis Wells & Wellington 1985 1890 Perochirus mestoni De Vis (Shea 1995) Dactyloperus mestoni Wells & Wellington 1985 1985 Dactyloperus kingi Wells & Wellington (Sistrom, Hutchinson, Hutchinson & Donnellan 2009) Remarks: The common names 'Northern Dtella' and 'Australian House Gecko' have also been used for australis. Previously regarded as a synonym of variegata (e.g., Kluge 1963; Wermuth 1965). Records from New Guinea refer to another species (Shea, Wilson, Greenlees, Amey & Hobson 2018). Distribution: Australia (New South Wales, Queensland [incl. Fraser Island, Torres Strait Islands]). Gehyra einasleighensis Einasleigh Rock Dtella Einasleigh-Vierkrallengecko Einasleigh-stumpfingergekko 2017 Gehyra einasleighensis Bourke, Pratt, Vanderduys & Moritz Distribution: Australia (Queensland). Gehyra electrum Amber Rock Dtella, Mount Surprise Rock Dtella Mount-Surprise-Vierkrallengecko Mount Surprise-stumpfingergekko 2019 Gehyra electrum Zozaya, Fenker & MacDonald Distribution: Australia (Queensland). Gehyra fehlmanni Kanchanaburi Four-clawed Gecko, (Fehlmann's Four-clawed Gecko) Fehlmanns Vierkrallengecko Kanchanaburi-stumpfingergekko 1962 Peropus fehlmanni Taylor Gehyra fehlmanni Wermuth 1965 Remarks: Although reported from Cambodia (e.g., Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as Gehyra cf. fehlmanni]; Nguyen, Ho & Nguyen 2009) and Vietnam (e.g., Bobrov 1995; Nguyen, Ho & Nguyen 2009), these records were not accepted subsequently and need confirmation (David & Ineich 2009). Distribution: Thailand. Gehyra fehlmanni © Henrik Bringsøe Gehyra fenestrula Hamersley Range Spotted Dtella Hamersley-Vierkrallengecko Hamersley-stumpfingergekko 2018 Gehyra fenestrula Doughty, Bauer, Pepper & Keogh Remarks: Previously regarded as a population of punctata (Doughty, Bauer, Pepper & Keogh 2018). Distribution: Australia (Western Australia). Gehyra finipunctata Small-spotted Mid-west Rock Dtella Mount-Minnie-Vierkrallengecko Mount Minnie-stumpfingergekko 2018 Gehyra punctulata Doughty, Bauer, Pepper & Keogh 2018 [not Phyria punctulata Gray 1842; nomen oblitum] 2018 Gehyra finipunctata Ellis, Doughty, Bauer, Pepper & Keogh [substitute name for Gehyra punctulata Doughty, Bauer, Pepper & Keogh 2018] Remarks: Previously regarded as a population of punctata (Doughty, Bauer, Pepper & Keogh 2018). Distribution: Australia (Western Australia). Gehyra gemina Plain Tree Dtella Einfarbiger Baum-Vierkrallengecko Ensfarvet Træ-stumpfingergekko 2020 Gehyra gemina Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz Distribution: Australia (Northern Territory, Western Australia). Gehyra georgpotthasti Western Giant Stump-toed Gecko, Vanuatu Giant Gecko Westlicher Riesen-Vierkrallengecko Vestlig Kæmpe-stumpfingergekko 2012 Gehyra georgpotthasti Flecks, Schmitz, Böhme, Henkel & Ineich Remarks: Previously considered to be populations of vorax. Introduced to French Polynesia (Tuamotu Archipelago: Fakarava). A record from Norfolk Island, Australia (Boulenger 1885) is presumed to be in error or an introduction, which did not establish a population (Cogger, cameron & Cogger 1983; Cogger 1996; Bauer & Sadlier 2000; Flecks, Schmitz, Böhme, Henkel & Ineich 2012). Introduced to: Australia (Norfolk Island), French Polynesia. Distribution: New Caledonia (Loyalty Islands), Vanuatu (incl. Espiritu Santo). Gehyra girloorloo Girloorloo Dtella, Kimberley Karst Gecko Girloorloo-Vierkrallengecko Girloorloo-stumpfingergekko 2016 Gehyra girloorloo Oliver, Bourke, Pratt, Doughty & Moritz Distribution: Australia (Western Australia). Gehyra granulum Lesser Kimberley Spotted Dtella, Kimberley Granular-toed Gecko Kleiner Gefleckter Vierkrallengecko Lille Plettet Stumpfingergekko 2018 Gehyra granulum Doughty, Bourke, Tedeschi, Pratt, Oliver, Palmer & Craig Remarks: Previously regarded as a population of nana. Distribution: Australia (Western Australia). Gehyra incognita Northern Pilbara Dtella, Northern Pilbara Cryptic Gehyra Nördlicher Pilbara-Vierkrallengecko Nordlig Pilbara-stumpfingergekko 2018 Gehyra incognita Kealley, Doughty, Pepper, Keogh, Hillyer & Huey Distribution: Australia (Western Australia). Gehyra intermedia Ryukyu Stump-toed Gecko Ryukyu-Vierkrallengecko Ryukyu-stumpfingergekko 1902 Gehyra intermedia Brown Peropus intermedius Chrapliwy, Smith & Grant 1961 Remarks: Status uncertain, as the type specimen is lost. Possibly a synonym of Gekko hokouensis, according to the Reptile Database (reptile-database.reptarium.cz, viewed 17 April, 2013). Distribution: Japan (Ryukyu Islands). Gehyra interstitialis Western New Guinean Four-clawed Gecko Westpapuanischer Vierkrallengecko Vestpapuansk Stumpfingergekko 1894 Gehyra interstitialis Oudemans Peropus interstitialis Barbour 1912 Distribution: Indonesia (Western New Guinea), Papua New Guinea (Eastern New Guinea). Gehyra ipsa Purnululu Dtella, Bungle Bungle Ranges Dtella Purnululu-Vierkrallengecko Purnululu-stumpfingergekko 2005 Gehyra koira ipsa Horner Gehyra ipsa Cogger 2014 Remarks: Previously included the population now assigned to chimera (Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz 2020). Distribution: Australia (Western Australia). Gehyra kimberleyi Robust Termitaria Dtella, Robust Termitaria Gecko, (Kimberley Dtella) Derby-Vierkrallengecko Derby-stumpfingergekko 1983 Gehyra kimberleyi Börner & Schüttler Dactyloperus kimberleyi Wells & Wellington 1984 Remarks: Status of this species has been disputed. Some authors have regarded the publication in which the species was described as not constituting a publication for the purposes of zoological nomenclature. It was not mentioned by Wilson & Swan (2010). Considered a valid species herein, following Bauer (1994) and Oliver, Bourke, Pratt, Doughty & Moritz (2016). Distribution: Australia (Western Australia). Gehyra koira Koira Dtella, (King's Rock Dtella) Koira-Vierkrallengecko Koira-stumpfingergekko 2005 Gehyra koira Horner Remarks: Previously included the populations now assigned to calcitectus, chimera, ipsa, and lapistola (Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz 2020). Distribution: Australia (Northern Territory, Western Australia). Gehyra lacerata Lacerated Four-clawed Gecko, Western Four-clawed Gecko, (Kanchanaburi Four-clawed Gecko) Westthailändischer Vierkrallengecko Vestthailandsk Stumpfingergekko 1962 Peropus laceratus Taylor Gehyra lacerata Wermuth 1965 Remarks: Presence in Vietnam needs confirmation (Pauwels, Kunya & Bauer 2008; David & Ineich 2009), although reported from the country by several sources (e.g., Bobrov 1995; Nguyen, Ho & Nguyen 2009). Distribution: Thailand. Gehyra lampei Madang Dtella Madang-Vierkrallengecko Madang-stumpfingergekko 1913 Gehyra lampei Andersson Remarks: Previously regarded as a synonym of papuana (e.g., Loveridge 1935; Wermuth 1965). Distribution: Papua New Guinea (Eastern New Guinea). Gehyra lapistola Litchfield Rock Dtella Litchfield Felsen-Vierkrallengecko Litchfield Klippe-stumpfingergekko 2020 Gehyra lapistola Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz Distribution: Australia (Northern Territory). Gehyra lauta Gulf Tree Dtella Queensland-Baum-Vierkrallengecko Queensland-træ-stumpfingergekko 2020 Gehyra lauta Oliver, Prasetya, Tedeschi, Fenker, Ellis, Doughty & Moritz Distribution: Australia (Northern Territory, Queensland). Gehyra lazelli Southern Rock Dtella, (Lazell's Dtella) Südaustralischer Vierkrallengecko Sydaustralsk Stumpfingergekko 1985 Dactyloperus lazelli Wells & Wellington Gehyra lazelli Sistrom, Hutchinson, Hutchinson & Donnellan 2009 1985 Dactyloperus annetteae Wells & Wellington (Sistrom, Hutchinson, Hutchinson & Donnellan 2009) Remarks: Previously synonymized with variegata (e.g., Kluge 1991; Bauer 1994; Shea & Sadlier 1999). Revalidated by Sistrom, Hutchinson, Hutchinson & Donnellan (2009). Distribution: Australia (New South Wales, South Australia). Gehyra leopoldi Doberai Dtella Doberai-Vierkrallengecko Doberai-stumpfingergekko 1930 Gehyra leopoldi Brongersma Peropus leopoldi Chrapliwy, Smith & Grant 1961 Distribution: Indonesia (Western New Guinea), Papua New Guinea (Eastern New Guinea). Gehyra macra Large Pilbara Rock Dtella Grosser Pilbara-Vierkrallengecko Stor Pilbara-stumpfingergekko 2018 Gehyra macra Doughty, Bauer, Pepper & Keogh Remarks: Previously regarded as a population of punctata (Doughty, Bauer, Pepper & Keogh 2018). Distribution: Australia (Western Australia). Gehyra marginata Giant Moluccan Dtella Molukkanischer Vierkrallengecko Molukkansk Stumpfingergekko 1887 Gehyra marginata Boulenger 1887 Gehyra fischeri Strauch (Bauer 1994) Remarks: Beckon (1992) tentatively considered marginata to be a synonym of vorax, but Flecks, Schmitz, Böhme, Henkel & Ineich (2012) showed that they are separate species. Distribution: Indonesia (Halmahera, Kai Islands, Morotai, Obi, Ternate). Gehyra media Medium Pilbara Spotted Rock Dtella Mittlerer Pilbara-Vierkrallengecko Middelstor Pilbara-stumpfingergekko 2018 Gehyra media Doughty, Bauer, Pepper & Keogh Remarks: Previously regarded as a population of punctata (Doughty, Bauer, Pepper & Keogh 2018). Distribution: Australia (Western Australia). Gehyra membranacruralis New Guinean Giant Stump-toed Gecko Grosser Papuanischer Vierkrallengecko Stor Papuansk Stumpfingergekko 1989 Gehyra membranacruralis King & Horner Remarks: Beckon (1992) considered membranacruralis to be a synonym of vorax, but it is now regarded as a valid species (e.g., Bauer 1994; Flecks, Schmitz, Böhme, Henkel & Ineich 2012). Distribution: Indonesia (Western New Guinea), Papua New Guinea (Eastern New Guinea, Louisiade Archipelago). Gehyra micra Small Pilbara Spotted Rock Dtella Kleiner Pilbara-Vierkrallengecko Lille Pilbara-stumpfingergekko 2018 Gehyra micra Doughty, Bauer, Pepper & Keogh Remarks: Previously regarded as a population of punctata (Doughty, Bauer, Pepper & Keogh 2018). Distribution: Australia (Western Australia). Gehyra minuta Dwarf Dtella Zwerg-Vierkrallengecko Dværg-stumpfingergekko 1982 Gehyra minuta King Dactyloperus minutus Wells & Wellington 1985 Distribution: Australia (Northern Territory). Gehyra montium Watarru Spotted Dtella, (Centralian Spotted Dtella, Centralian Dtella) Watarru-Vierkrallengecko Watarru-stumpfingergekko 1982 Gehyra montium Storr Dactyloperus montium Wells & Wellington 1984 Distribution: Australia (Northern Territory, South Australia, Western Australia). Gehyra moritzi Corroboree Rock Spotted Dtella Corroboree-Vierkrallengecko Corroboree-stumpfingergekko 2014 Gehyra moritzi Hutchinson, Sistrom, Donnellan & Hutchinson Distribution: Australia (Northern Territory). Gehyra multiporosa Mitchell Plateau Dtella, Multi-pored Dtella Vielporiger Vierkrallengecko Mangeporet Stumpfingergekko 2012 Gehyra multiporosa Doughty, Palmer, Sistrom, Bauer & Donnellan Distribution: Australia (Western Australia). Gehyra mutilata Common Four-clawed Gecko, Stump-toed Gecko, Stump-tailed Gecko, Four-clawed House Gecko, Eastern House Gecko, Tender-skinned House Gecko, Skin-shedding Dtella, Buff House Gecko, Mutilating Gecko, (Pacific Gecko) Gewöhnlicher Vierkrallengecko, Pazifikgecko Almindelig Stumpfingergekko 1835 Hemidactylus mutilatus Wiegmann Peropus mutilatus Fitzinger 1843 Peripia mutilata Günther 1873 Gehyra mutilata Boulenger 1885 1836 Hemidactylus peronii Duméril & Bibron (Bauer 1994) Gehyra peronii Wells & Wellington 1985 1857 Dactyloperus insulensis Girard (Bauer 1994) Gehyra insulensis Wells & Wellington 1985 1865 Gecko pardus Tytler (Bauer 1994) Gehyra pardus Wells & Wellington 1985 1869 Peropus packardii Cope (Bauer 1994) Gehyra packardii Wells & Wellington 1985 1884 Hemidactylus navarri Dugès (Bauer 1994) 1900 Gehyra butleri Boulenger (Grismer 2011) Peropus butleri Smith 1930 1913 Gehyra beebei Annandale (Bauer 1994) 1985 Gehyra harrieti Wells & Wellington (Bauer 1994) Other common names: butleri: Butler's Four-clawed Gecko insulensis: Pacific Stump-toed Gecko, Pacific Gecko Remarks: Rocha, Ineich & Harris (2009) suggested that insulensis might represent a valid species, comprising the Pacific populations of mutilata, however, they did not make a formal revalidation, pending further studies. Nevertheless, Flecks, Schmitz, Böhme, Henkel & Ineich (2012); Zug, Ineich, Pregill & Hamilton (2012), and Zug (2013) listed insulensis as a separate species. According to Zhao & Adler (1993), erroneously recorded as introduced to New Zealand (e.g., Robb 1980). Not listed for Chile (Easter Island) by Gamboa (2020). Introduced to: Christmas Island, French Guiana, India (Andaman Islands, Uttar Pradesh), Indonesia (Talaud Islands), Mauritius (incl. Rodrigues), Mexico (Baja California Sur, Chiapas, Jalisco, Nayarit, San Luis Potosi, Sinaloa), New Zealand, Reunion, USA (California, Florida, Hawaii), Vanuatu. Distribution: American Samoa, Brunei, Cambodia, China (Guangdong, Hainan, Hong Kong, Yunnan), Clipperton, Cocos (Keeling) Islands, Cook Islands (incl. Rarotonga), East Timor (incl. Atauro Island), Easter Island, Fiji (incl. Rotuma), French Polynesia, Guam, India (Andaman & Nicobar Islands [Andaman Islands, Nicobar Islands], Kerala), Indonesia (Alor, Ambon, Anambas Islands, Aru Islands, Bacan, Bali, Banda Islands, Banggai Islands, Bangka, Batu Islands, Buru, Buton, Ende, Enggano, Flores, Gorong Archipelago, Halmahera, Hoga, Java, Kabaena, Kai Islands, Kalimantan, Kangean Islands, Karimunjawa Islands, Komodo, Krakatau, Legundi, Lombok, Morotai, Natuna Islands, Nias, Panaitan, Penida, Rote, Sangir Islands [incl. Biaro], Saparua, Sebesi, Selayar, Sula Islands, Sulawesi, Sumatra, Sumba, Sumbawa, Tanahjampea, Tanimbar Islands, Ternate, Tinjil, Waigeo, West Timor, Western New Guinea, Wetar), Japan (Ryukyu Islands [Okinawa Islands, Sakishima Islands, Satsunan Islands]), Kiribati, Laos, Madagascar (incl. Nosy Be), Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Penang Island, Perhentian Islands, Seribuat Archipelago (incl. Aur, Pemanggil, Tioman)]), Marshall Islands (incl. Arno Atoll), Micronesia (incl. Pohnpei [incl. Ant Atoll]), Myanmar, Nauru, Northern Mariana Islands, Palau, Papua New Guinea (incl. Admiralty Islands, Bismarck Archipelago [New Britain, New Ireland, Umboi], d'Entrecasteaux Islands, Eastern New Guinea, Louisiade Archipelago [Rossel, Sudest], Woodlark, North Solomon Islands [Bougainville]), Philippines (Apo, Babuyan Islands [Calayan, Camiguin Norte, Dalupiri, Fuga], Balabac, Balicasag, Bantayan, Batan Islands [Batan], Bohol, Bonoon, Boracay, Calagnaan, Calamian Islands [Calauit], Caluya, Camiguin Sur, Camotes Islands [Pacijan, Ponson, Poro], Carabao, Catanduanes, Cebu, Clara, Danjugan, Dinagat, Duitay, Dumaran, Gigante Islands, Guimaras, Inampulugan, Jao, Lapinig Islands [Lapinig Chico, Lapinig Grande], Leyte, Luzon, Mactan, Mantique, Marinduque, Mindanao, Mindoro, Nadulao, Negros, Palawan, Pan de Azucar, Panay, Panubulon, Samar, Sibuyan, Sicogon, Siquijor, Sulu Islands [Bongao, Bubuan, Jolo], Tablas, Tilmubo), Pitcairn Islands, Seychelles (incl. Aldabra, Praslin), Singapore, Solomon Islands (incl. Guadalcanal, Santa Cruz Islands), Sri Lanka, Taiwan, Thailand, Tokelau, Tonga, Vietnam, Wake, Western Samoa. Gehyra mutilata © Rune Midtgaard Gehyra nana Northern Spotted Dtella, Northern Spotted Rock Gecko Nördlicher Gefleckter Vierkrallengecko Nordlig Plettet Stumpfingergekko 1978 Gehyra nana Storr Dactyloperus nana Wells & Wellington 1984 Remarks: Doughty, Bourke, Tedeschi, Pratt, Oliver, Palmer & Craig (2018) did not mention the occurrence of this species in Queensland, neither did they include the state in their distribution map. It is unclear to me whether Queensland records (e.g., Cogger 2014; Vanderduys, Hines, Gynther, Kutt & Absolon 2011; Wilson & Swan 2010; Wilson 2005) are still valid or have been reassigned to other species. Distribution: Australia (Northern Territory, Queensland, Western Australia). Gehyra nana © Rune Midtgaard Gehyra occidentalis West Kimberley Dtella, Kimberley Plateau Dtella Westlicher Kimberley-Vierkrallengecko Vestlig Kimberley-stumpfingergekko 1984 Gehyra occidentalis King Dactyloperus occidentalis Wells & Wellington 1985 Phyriadoria occidentalis Wells & Wellington 1989 Distribution: Australia (Western Australia). Gehyra oceanica Oceanic Gecko Pazifischer Vierkrallengecko Pacifisk Stumpfingergekko 1830 Gecko oceanicus Lesson Gehyra oceanica Gray 1845 Hemidactylus oceanicus Schmeltz 1874 Peropus oceanicus Barbour 1912 1820 Stellio argyropus Tilesius von Tilenau (Bauer 1994) 1834 Gehyra pacifica Gray (Bauer 1994) 1836 Hemidactylus oualensis Duméril & Bibron (Bauer 1994) 1842 Gehyra gularis Gray (Bauer 1994) Remarks: Presumed extirpated in Guam (Bauer 1994). Presence in mainland New Guinea needs confirmation, according to Beckon (1992), who believed the species to be rare or absent there. Nevertheless, it has been listed from the island by many subsequent publications (e.g., Bauer 1994; Rösler, Glaw & Günther 2005; Kraus & Allison 2006). Records from the Loyalty Islands (New Caledonia) (Burt & Burt 1932) refer to vorax (Bauer & Sadlier 2000). Records from the Isle of Pines, off the New Caledonian mainland, are based on wrong locality data (Bauer & Sadlier 2000). Records from Lord Howe & Norfolk Islands (Australia) are erroneous (Cogger 1996). Records from the Australian mainland (incl. Torres Strait Islands) are doubtful (Bauer 1994; Cogger 1996). Not listed for Chile (Easter Island) by Gamboa (2020). Extinct in: Guam. Introduced to: New Zealand, USA (Hawaii). Distribution: American Samoa, Cook Islands (incl. Rarotonga), Easter Island, Fiji (incl. Rotuma), French Polynesia, Indonesia (Halmahera, Kai Islands, Morotai, Supiori, Ternate, Western New Guinea), Kiribati, Marshall Islands (incl. Arno Atoll), Micronesia (incl. Pohnpei [incl. Ant Atoll]), Nauru, Niue, Northern Mariana Islands, Palau, Papua New Guinea (incl. Admiralty Islands [Manus], Bismarck Archipelago [New Britain, New Ireland], d'Entrecasteaux Islands [Fergusson, Normanby], Eastern New Guinea, Louisiade Archipelago [Rossel, Sudest], Woodlark, North Solomon Islands [Bougainville]), Solomon Islands (incl. Bellona, Taumako, Guadalcanal, Rennell, Santa Cruz Islands), Tokelau, Tonga, Tuvalu, Vanuatu (incl. Espiritu Santo), Wallis & Futuna, Western Samoa. Gehyra oceanica © Rune Midtgaard Gehyra ocellata Pilbara Island Dtella, Pilbara Island Gehyra Pilbara-Inseln-Vierkrallengecko Barrow Island-stumpfingergekko 2018 Gehyra ocellata Kealley, Doughty, Pepper, Keogh, Hillyer & Huey Distribution: Australia (Western Australia). Gehyra pamela Arnhem Land Spotted Dtella, Arnhem Land Watercourse Dtella, Arnhem Land Spotted Gecko Arnhemland-Vierkrallengecko Arnhemland-stumpfingergekko 1982 Gehyra pamela King Dactyloperus pamela Wells & Wellington 1984 Phyriadoria pamela Wells & Wellington 1989 Distribution: Australia (Northern Territory). Gehyra papuana Papuan Dtella Papuanischer Vierkrallengecko Papuansk Stumpfingergekko 1874 Gehyra papuana Meyer Peropus papuanus Chrapliwy, Smith & Grant 1961 Remarks: May be present in Belau and Marianas Islands from where it has been recorded by Dryden & Taylor (1969), although at least the Belau specimens are probably referable to brevipalmata (Bauer 1994). Distribution: Indonesia (Western New Guinea), Papua New Guinea (Eastern New Guinea). Gehyra paranana Northwestern Spotted Dtella Nordwestlicher Gefleckter Vierkrallengecko Nordvestlig Plettet Stumpfingergekko 2018 Gehyra paranana Doughty, Bourke, Tedeschi, Pratt, Oliver, Palmer & Craig Remarks: Previously regarded as a population of nana. Distribution: Australia (Northern Territory). Gehyra peninsularis Burrup Peninsula Rock Dtella Burrup-Vierkrallengecko Burrup-stumpfingergekko 2018 Gehyra peninsularis Doughty, Bauer, Pepper & Keogh Remarks: Previously regarded as a population of punctata (Doughty, Bauer, Pepper & Keogh 2018). Distribution: Australia (Western Australia). Gehyra pilbara Common Pilbara Dtella, (Pilbara Gecko) Gewöhnlicher Pilbara-Vierkrallengecko Almindelig Pilbara-stumpfingergekko 1965 Gehyra pilbara Mitchell Dactyloperus pilbara Wells & Wellington 1984 1982 Gehyra cognata Börner & Schüttler (Bauer 1994) Dactyloperus cognata Wells & Wellington 1985 Distribution: Australia (Northern Territory, Western Australia). Gehyra pluraporosa North Kimberley Spotted Dtella, Northern Kimberley Gecko Nördlicher Kimberley-Vierkrallengecko Nordlig Kimberley-stumpfingergekko 2018 Gehyra pluraporosa Doughty, Bourke, Tedeschi, Pratt, Oliver, Palmer & Craig Remarks: Previously regarded as a population of nana. Distribution: Australia (Western Australia). Gehyra polka Large-spotted Mid-west Rock Dtella Noondie-Vierkrallengecko Noondie-stumpfingergekko 2018 Gehyra polka Doughty, Bauer, Pepper & Keogh Remarks: Previously regarded as a population of punctata (Doughty, Bauer, Pepper & Keogh 2018). Distribution: Australia (Western Australia). Gehyra pseudopunctata Southern Kimberley Spotted Dtella, Southern Kimberley Spotted Gecko Südlicher Kimberley-Vierkrallengecko Sydlig Kimberley-stumpfingergekko 2018 Gehyra pseudopunctata Doughty, Bourke, Tedeschi, Pratt, Oliver, Palmer & Craig Remarks: Previously regarded as a population of nana. Distribution: Australia (Western Australia). Gehyra pulingka Umuwa Spotted Dtella Umuwa-Vierkrallengecko Umuwa-stumpfingergekko 2014 Gehyra pulingka Hutchinson, Sistrom, Donnellan & Hutchinson Distribution: Australia (Northern Territory, South Australia, Western Australia). Gehyra punctata Spotted Pilbara Rock Dtella, (Western Spotted Dtella, Spotted Rock Dtella, Spotted Gecko) Gefleckter Pilbara-Vierkrallengecko Plettet Pilbara-stumpfingergekko 1914 Peropus variegatus var. punctatus Fry Gehyra variegata punctata Mitchell 1955 Gehyra punctata Worrell 1963 Dactyloperus punctata Wells & Wellington 1984 1965 Gehyra fenestra Mitchell (Storr 1982; Doughty, Bauer, Pepper & Keogh 2018) Dactyloperus fenestra Wells & Wellington 1985 Remarks: Validity as a separate species confirmed by Bustard (1965b). Previously included the populations now assigned to fenestrula, macra, media, micra, peninsularis, polka, and punctulata (Doughty, Bauer, Pepper & Keogh 2018). Distribution: Australia (Western Australia). Gehyra purpurascens Purple Arid Dtella, Purple Dtella, Purplish Dtella Purpur-Vierkrallengecko Purpur-stumpfingergekko 1982 Gehyra purpurascens Storr Dactyloperus purpurascens Wells & Wellington 1984 Distribution: Australia (Northern Territory, Queensland, South Australia, Western Australia). Gehyra robusta Robust Dtella Dicker Vierkrallengecko Tyk Stumpfingergekko 1983 Gehyra robusta King Dactyloperus robustus Wells & Wellington 1984 Distribution: Australia (Queensland). Gehyra rohan Rohan Stump-toed Gecko Rohan-Vierkrallengecko Rohan-stumpfingergekko 2016 Gehyra rohan Oliver, Clegg, Fisher, Richards, Taylor & Jocque Distribution: Papua New Guinea (Bismarck Archipelago [Admiralty Islands (Los Negros Island, Manus)], Saint Matthias Islands [Mussau Island]). Gehyra serraticauda Saw-tailed Dtella Sägeschwanz-Vierkrallengecko Savhalet Stumpfingergekko 2014 Gehyra serraticauda Skipwith & Oliver Distribution: Indonesia (Western New Guinea). Gehyra spheniscus Lesser Wedge-toed Dtella, Small Wedge-toed Dtella Kleiner Vierkrallengecko Lille Stumpfingergekko 2012 Gehyra spheniscus Doughty, Palmer, Sistrom, Bauer & Donnellan Distribution: Australia (Western Australia). Gehyra unguiculata Crescent-marked Dtella, Crescent-marked Pilbara Dtella Halbmond-Vierkrallengecko Halvmåne-stumpfingergekko 2018 Gehyra unguiculata Kealley, Doughty, Pepper, Keogh, Hillyer & Huey Distribution: Australia (Western Australia). Gehyra variegata Southwestern Variegated Dtella, (Variegated Dtella, Common Dtella, Tree Dtella, Tree Gecko) Südwestlicher Bunter Vierkrallengecko Sydvestlig Broget Stumpfingergekko 1836 Hemidactylus variegatus Dumeril & Bibron Peropus variegatus Fitzinger 1843 Peripia variegata Gray 1867 Gehyra variegata Boulenger 1885 Dactyloperus variegatus Wells & Wellington 1984 Remarks: Previously included lazelli and the populations now assigned to capensis, crypta, incognita, ocellata, unguiculata, and versicolor. New Guinean records (e.g., Rooij 1915; Whitaker, Whitaker & Mills 1982) must refer to other species (Bauer 1994). A record (Waite 1927) from Kangaroo Island, South Australia, is regarded as erroneous, since it has not been mentioned or mapped from the island in subsequent works (e.g., Hutchinson 2010). Distribution: Australia (South Australia, Western Australia). Gehyra versicolor Eastern Variegated Dtella Östlicher Bunter Vierkrallengecko Østlig Broget Stumpfingergekko 2014 Gehyra versicolor Hutchinson, Sistrom, Donnellan & Hutchinson Remarks: Consists of populations previously assigned to variegata. Distribution: Australia (New South Wales, Northern Territory, Queensland, South Australia, Victoria). Gehyra vorax Eastern Giant Stump-toed Gecko, Voracious Gecko, Giant Forest Gecko, Fiji Giant Gecko Östlicher Riesen-Vierkrallengecko Østlig Kæmpe-stumpfingergekko 1858 Gehyra vorax Girard Remarks: Previously regarded as a synonym of oceanica (e.g., de Rooij 1915; Wermuth 1965). Revalidated as a separate species by Beckon (1992). Now restricted to Fiji. Neither Beckon (1992), nor Flecks, Schmitz, Böhme, Henkel & Ineich (2012) examined any specimens from Tonga, although they tentatively included the country in the distribution, however, no verified specimens from Tonga exist (Zug, Ineich, Pregill & Hamilton 2012). Records from Loyalty Islands, Vanuatu, and Norfolk Island refer to georgpotthasti. Records from the Moluccas refer to marginata. Records from New Guinea and satellites refer to marginata or membranacruralis. Introduced to French Polynesia (Flecks, Schmitz, Böhme, Henkel & Ineich 2012). Introduced to: French Polynesia. Distribution: Fiji. Gehyra wongchan Lopburi Dtella Lopburi-Vierkrallengecko Lopburi-stumpfingergekko 2022 Gehyra wongchan Pauwels, Meesok, Kunya, Donbundit & Sumontha Distribution: Thailand. Gehyra xenopus Crocodile-faced Dtella Stülpnasen-Vierkrallengecko Braknæset Stumpfingergekko 1978 Gehyra xenopus Storr Dactyloperus xenopus Wells & Wellington 1984 Phyriadoria xenopus Wells & Wellington 1989 Distribution: Australia (Western Australia).