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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update: December 31st, 2022.


Taxonomy of the family Phyllodactylidae
Bibliography of the genus Ptyodactylus
Biodiversity of the family Phyllodactylidae








Genus
Ptyodactylus

Fan-toed Geckos, Fan-footed Geckos, Lobe-footed Geckos, Fan-fingered Geckos

Fächerfingergeckos, Fächerfussgeckos

Viftefingergekkoer

1820 Ptyodactylus Goldfuss (type species: Gecko lobatus Geoffroy 1809)
Contents: 13 species, of which 6 (46.2%) are endemic.
Endemism: 0% 100%
Remarks: A single record of the genus Ptyodactylus from Iran refer to Asaccus nasrullahi (Werner 2006).
Distribution: N. Africa, Middle East, Pakistan.
Reported from: Algeria, Benin, Burkina Faso, Cameroon, Chad, Djibouti, Egypt (incl. Sinai), Eritrea, Ethiopia, Ghana, Iraq, Israel, Ivory Coast, Jordan, Lebanon, Libya, Mali, Mauritania, Morocco, Niger, Nigeria, Oman, Pakistan, Saudi Arabia (incl. Farasan Islands), Somalia, Sudan, Syria, Togo, United Arab Emirates, Yemen.

Ptyodactylus ananjevae

Southern Jordan Fan-toed Gecko

Südjordanischer Fächerfingergecko

Sydjordansk Viftefingergekko

2013 Ptyodactylus ananjevae Nazarov, Melnikov & Melnikova

Distribution: Jordan.


Ptyodactylus dhofarensis

Dhofar Fan-toed Gecko

Dhofar-Fächerfingergecko

Dhofar-viftefingergekko

2013 Ptyodactylus dhofarensis Nazarov, Melnikov & Melnikova

Distribution: Oman.


Ptyodactylus guttatus

Spotted Fan-toed Gecko, Sinai Fan-toed Gecko

Gefleckter Fächerfingergecko

Plettet Viftefingergekko

1827 Ptyodactylus guttatus Heyden
Ptyodactylus hasselquistii guttatus Welch 1983
1834 Gecko maculatus Schinz [1833-35] [substitute name for Ptyodactylus guttatus Heyden 1827] (Heimes 1987)
1892 Ptyodactylus lacazzi Boutan (Heimes 1987)

Remarks: Previously regarded as a synonym of hasselquistii (e.g., Loveridge 1947; Wermuth 1965). Revalidated by Heimes (1987). Previously included siphonorhina. A record from Lebanon (Werner 1939) refer to puiseuxi (Hraoui-Bloquet, Sadek, Sindaco & Venchi 2002). Werner & Sivan (1994) considered Oman populations of hasselquisti as guttatus. Records from Saudi Arabia (e.g., Sindaco & Jeremchenko 2008) and Sudan (e.g., Heimes 1987; Berre 1989) are regarded as doubtful (Aloufi, Amr, Baker & Hamidan 2019; Al-Johany Amr Baha-El-Din Disi Werner 2021).
Distribution: Egypt (incl. Sinai), Israel, Jordan, Syria.


Ptyodactylus hasselquistii

Egyptian Fan-toed Gecko, Yellow Fan-toed Gecko, (Hasselquist's Fan-toed Gecko)

Ägyptischer Fächerfingergecko

Egyptisk Viftefingergekko

1798 Lacerta hasselquistii Donndorff
Ptyodactylus hasselquistii Duméril & Bibron 1836
1809 Gecko lobatus Geoffroy (Heimes 1987)
Ptyodactylus lobatus Boulenger 1885
1820 Gekko ascalabotes Merrem (Heimes 1987)
1829 Stenodactylus guttatus Cuvier (Metallinou & Crochet 2013)
1995 Ptyodactylus hasselquistii krameri Werner

Remarks: Previously reported from Iran (e.g., Schmidt 1955; Anderson 1963, 1968, 1974; Schleich 1977), but records of Ptyodactylus from this country are now referred to Asaccus nasrullahi (Werner 2006). Presence in Lebanon has been disputed (Bosch 1998). Tamar & Moravec (2022) documented the species from the country, although probably based on a recent introduction. Hraoui-Bloquet's (1981) record of hasselquistii from Lebanon refers to puiseuxi (Hraoui-Bloquet, Sadek, Sindaco & Venchi 2002). The species has also been reported from Syria (e.g., Arnold 1986; Disi & Böhme 1996), but in error, probably due to confusion with other species of the genus (Martens 1997). Oman and United Arab Emirates populations refer to dhofarensis, orlovi, and ruusaljibalicus (Nazarov, Melnikov & Melnikova 2013; Simó-Riudalbas, Metallinou, Pous, Els, Jayasinghe, Péntek-Zakar, Wilms, Al-Saadi & Carranza 2017; Carranza, Els & Burriel-Carranza 2021). Records from Iraq (e.g., Leviton, Anderson, Adler & Minton 1992; Afrasiab & Mohamad 2009; Amr 2009; Al-Barazengy, Salman & Hameed 2015; Mohammed, Rhadi, Rastegar-Pouyani, Rastegar-Pouyani & Yousefkhani 2017) are regarded as doubtful (Heimes 1987; Els, Aloufi, Al-Johany, Amr, Baha-El-Din, Disi & Werner 2021).
Distribution: Egypt (incl. Sinai), Israel, Jordan, Saudi Arabia (incl. Farasan Islands), Sudan, Yemen.


Ptyodactylus homolepis

Pakistani Fan-toed Gecko, Sind Fan-toed Gecko

Pakistanischer Fächerfingergecko

Pakistansk Viftefingergekko

1876 Ptyodactylus homolepis Blanford

Remarks: Records of homolepis from Socotra (e.g., Loveridge 1947; Wermuth 1965) refer to Ptyodactylus socotranus Steindachner 1902, regarded as a subspecies of homolepis by both firstmentioned authors, but shown to be a synonym of Haemodracon riebecki by Eiselt (1962). Sharma (1971) and Sahi & Duda (1985) erroneously recorded homolepis from Jammu & Kashmir, India (Venugopal 2010).
Distribution: Pakistan.


Ptyodactylus orlovi

Eastern Arabian Fan-toed Gecko

Ostarabischer Fächerfingergecko

Østarabisk Viftefingergekko

2013 Ptyodactylus orlovi Nazarov, Melnikov & Melnikova

Distribution: Oman, United Arab Emirates.


Ptyodactylus oudrii

Moroccan Fan-toed Gecko, (Oudri's Fan-toed Gecko)

Marokkanischer Fächerfingergecko

Marokkansk Viftefingergekko

1880 Ptyodactylus oudrii Lataste
Ptyodactylus hasselquistii oudrii Anderson 1898

Remarks: Revalidated as a separate species by Heimes (1987). Validity confirmed by Perera & Harris (2010) who suggested that it is a species complex.
Distribution: Algeria, Morocco.


Ptyodactylus puiseuxi

Levant Fan-toed Gecko, Israeli Fan-toed Gecko

Levantischer Fächerfingergecko

Levantisk Viftefingergekko

1893 Ptyodactylus puiseuxi Boutan
Ptyodactylus hasselquistii puiseuxi Loveridge 1947
1893 Ptyodactylus barroisi Boutan (Heimes 1987)
1893 Ptyodactylus bischoffsheimi Boutan (Heimes 1987)
1893 Ptyodactylus montmahoui Boutan (Heimes 1987)
1894 Ptyodactylus lobatus syriacus Peracca (Heimes 1987)
1914 Ptyodactylus lobatus sancti-montis Barbour (Heimes 1987)

Remarks: Previously regarded as a synonym of hasselquistii (e.g., Wermuth 1965). Revalidated by Heimes (1987). Although reported by several authors from Iraq (e.g., Leviton, Anderson, Adler & Minton 1992), Afrasiab & Mohamad (2009) were the first to provide a record with locality.
Distribution: Iraq, Israel, Jordan, Lebanon, Saudi Arabia, Syria.


Ptyodactylus ragazzii

Eritrean Fan-toed Gecko, (Sahelian Fan-toed Gecko, Ragazzi's Fan-toed Gecko)

Eritreische Fächerfingergecko

Eritreisk Viftefingergekko

1898 Ptyodactylus hasselquistii var. ragazzii Anderson
Ptyodactylus ragazzii Schleich, Kästle & Kabisch 1996

Remarks: Now restricted to Northeast Africa. Previously included togoensis and the populations now assigned to rivapadialorum (Trape 2017). Records from West Africa refer to these species (Metallinou, Cervenka, Crochet, Kratochvil, Wilms, Geniez, Shobrak, Brito & Carranza 2015; Trape 2017). Baha el Din (1999) presented evidence establishing ragazzi and hasselquistii as separate species, based on sympatry in southeastern Egypt. Already Schleich, Kästle & Kabisch (1996) and Smith, Robinson, Chiszar & Breukelen (1998) treated ragazzi as a distinct species, but without discussion. Records from Sudan (e.g., Heimes 1987; Schleich, Kastle & Kabisch 1996) are provisionally treated as valid herein, but confirmation is needed (Spawls, Mazuch & Mohammad 2023).
Distribution: Djibouti, Egypt, Eritrea, Ethiopia, Somalia, Sudan.


Ptyodactylus rivapadialorum

Mauritanian Fan-toed Gecko, (Riva and Padial’s Fan-footed Gecko)

Mauritanischer Fächerfingergecko

Mauritansk Viftefingergekko

2017 Ptyodactylus rivapadiali Trape
Ptyodactylus rivapadialorum [justified emendation] Midtgaard 2019

Remarks: Previously regarded as a population of ragazzii, or togoensis (e.g., Werner 1919; Bauer, Tchibozo, Pauwels & Lenglet 2006; Baha el Din 2006). The original specific epithet, rivapadiali, was corrected from singular to plural, rivapadialorum, to comply with Latin grammar, because it is a constructed name honouring two people, Ignacio de la Riva and José Padial (Midtgaard 2019).
Distribution: Mauritania.





Ptyodactylus ruusaljibalicus

Ruus al Jibal Fan-toed Gecko

Ruus-al-Jibal-Fächerfingergecko

Ruus al Jibal-viftefingergekko

2017 Ptyodactylus ruusaljibalicus Simó-Riudalbas, Metallinou, Pous, Els, Jayasinghe, Péntek-Zakar, Wilms, Al-Saadi & Carranza

Distribution: Oman, United Arab Emirates.


Ptyodactylus siphonorhina

Northeast Saharan Fan-toed Gecko, (Sahara Fan-toed Gecko)

Nordostsaharischer Fächerfingergecko

Nordøstsaharisk Viftefingergekko

1896 Ptyodactylus hasselquistii var. siphonorhina Anderson
Ptyodactylus siphonorhina Baha el Din 2006

Remarks: Previously regarded as a synonym of guttatus (e.g., Heimes 1987; Werner & Sivan 1994). Tentatively regarded as a separate species by Baha el Din (2006). Validity confirmed by Metallinou, Cervenka, Crochet, Kratochvil, Wilms, Geniez, Shobrak, Brito & Carranza (2015).
Distribution: Egypt.


Ptyodactylus togoensis

Sahelian Fan-toed Gecko

Sahel-Fächerfingergecko

Sahel-viftefingergekko

1901 Ptyodactylus hasselquisti var. togoensis Tornier
Ptyodactylus togoensis Metallinou, Cervenka, Crochet, Kratochvil, Wilms, Geniez, Shobrak, Brito & Carranza 2015

Remarks: Previously regarded as a synonym of ragazzii (e.g., Werner 1919; Bauer, Tchibozo, Pauwels & Lenglet 2006; Baha el Din 2006). Revalidated by Metallinou, Cervenka, Crochet, Kratochvil, Wilms, Geniez, Shobrak, Brito & Carranza (2015). Records from Mauritania refer to rivapadialorum (Trape 2017).
Distribution: Algeria, Benin, Burkina Faso, Cameroon, Chad, Ghana, Ivory Coast, Libya, Mali, Niger, Nigeria, Togo.