Reptiles of
North Macedonia [formerly FYR Macedonia]

 

Bibliography of the reptiles of North Macedonia Taxonomic history of the reptiles of North Macedonia Introduced species
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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist
General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial.
Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:
Lizards (Sauria)
= endemicEX = extinct or assumed extinct

Species
References
 Family Anguidae (Slowworms, etc.)
Genus Anguis (Slowworms)
 
Anguis colchica

Eastern Slowworm

Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010 Pozzi 1966 [as A. fragilis] Sterijovski, Tomovic & Ajtic 2014 [as A. fragilis] Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 [as A. fragilis]
 
Anguis graeca

Greek Slowworm

Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018
Genus Pseudopus (Eurasian Glass Lizard)
 
Pseudopus apodus

Eurasian Glass Lizard

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 Family Gekkonidae (Typical Geckos)
Genus Mediodactylus (Mediterranean Bent-toed Geckos)
 
Mediodactylus kotschyi

Balkan Bent-toed Gecko

Arnold & Ovenden 2002 Beutler in Boehme (ed.) 1981 Brelih & Dzukic 1974 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 Family Lacertidae (Typical Lizards)
Genus Algyroides (Keeled Lizards)
 
Algyroides nigropunctatus

Blue-throated Keeled Lizard

Arnold & Ovenden 2002 Bischoff in Boehme (ed.) 1981 Brelih & Dzukic 1974 Dimovski 1964 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Lacerta (Eurasian Green Lizards)
 
Lacerta agilis

Northern Eurasian Sand Lizard

Bischoff in Boehme (ed.) 1984 Brelih & Dzukic 1974 Dimovski 1959 Dimovski 1964 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 
Lacerta trilineata

Western Balkan Green Lizard

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Kornilios, Thanou, Lymberakis, Ilgaz, Kumlutas & Leache 2020 Nettmann & Rykena in Boehme (ed.) 1984 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 
Lacerta viridis

Eastern Green Lizard

Boehme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Dimovski 1959 Marzahn, Mayer, Joger, Ilgaz, Jablonski, Kindler, Kumlutas, Nistri, Schneewiess, Vamberger, Zagar & Fritz 2015 Necas, Modry & Zavadil 1997 Pafilis & Maragou 2013 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 [as "viridis complex"]
Genus Podarcis (Wall Lizards)
 
Podarcis erhardii

Aegean Wall Lizard

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Gruber in Boehme (ed.) 1986 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 
Podarcis muralis

Common European Wall Lizard

Brelih & Dzukic 1974 Dimovski 1959 Gruschwitz & Boehme in Boehme (ed.) 1986 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 
Podarcis tauricus

Balkan Wall Lizard

Brelih & Dzukic 1974 Dimovski 1959 Pozzi 1966 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Zootoca (Eurasian Forest Lizards)
 
Zootoca vivipara

Viviparous Lizard

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Dimovski 1964 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 Family Scincidae (Skinks)
Genus Ablepharus (Eurasian Snake-eyed Skinks)
 
Ablepharus kitaibelii

European Snake-eyed Skink

Brelih & Dzukic 1974 Dimovski 1959 Pozzi 1966 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016

Snakes (Serpentes)
= endemicEX = extinct or assumed extinct

Species
References
 Family Boidae (Typical Boas)
Genus Eryx (Sand Boas)
 
Eryx jaculus

Western Sand Boa

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Tokar & Obst in Boehme (ed.) 1993 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 Family Colubridae (Typical Snakes or Colubrids)
Genus Coronella (Palearctic Smooth Snakes)
 
Coronella austriaca

Northern Smooth Snake

Brelih & Dzukic 1974 Engelmann in Boehme (ed.) 1993 Geniez 2018 Jablonski, Nagy, Avci, Olgun, Kukushkin, Safaei-Mahroo & Jandzik 2019 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Dolichophis (Greater Whip Snakes)
 
Dolichophis caspius

Caspian Whip Snake

Brelih & Dzukic 1974 Geniez 2018 Nagy, Bellaagh, Wink, Paunovic & Korsos 2010 Pozzi 1966 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Elaphe (Palearctic-Oriental Rat Snakes)
 
Elaphe quatuorlineata

Western Four-lined Rat Snake

Arnold & Ovenden 2002 Boehme & Szczerbak in Boehme (ed.) 1993 Brelih & Dzukic 1974 Geniez 2018 Qi, Shi, Ma, Gao, Bu, Grismer, Li & Wang 2021 Schulz 1996 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Hierophis (Eurasian Whip Snakes)
 
Hierophis gemonensis

Balkan Whip Snake

Geniez 2018 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Natrix (Eurasian Keelbacks)
 
Natrix natrix

Common European Grass Snake

Brelih & Dzukic 1974 Geniez 2018 Kabisch in Boehme (ed.) 1999 Kindler, Boehme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroky & Fritz 2013 Pozzi 1966 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 
Natrix tessellata

Dice Snake

Brelih & Dzukic 1974 Geniez 2018 Necas, Modry & Zavadil 1997 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Ajtic, Tomovic, Djordjevic, Djurakic, Golubovic, Crnobrnja-Isailovic, Ballouard, Desmont, Groumpf & Bonnet 2011 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Platyceps (Flat-headed Whip Snakes)
 
Platyceps najadum

European Slender Whip Snake

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Darevsky & Szczerbak in Boehme (ed.) 1993 Geniez 2018 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Telescopus (Afro-Asian Cat Snakes)
 
Telescopus fallax

European Cat Snake

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1999 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
Genus Zamenis (Western Palearctic Rat Snakes)
 
Zamenis longissimus

Common Aesculapian Snake

Brelih & Dzukic 1974 Geniez 2018 Necas, Modry & Zavadil 1997 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 
Zamenis situla

Leopard Snake

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Obst, Szczerbak & Boehme in Boehme (ed.) 1993 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 Family Lamprophiidae (House Snakes, etc.)
Genus Malpolon (Montpellier Snakes)
 
Malpolon insignitus

Eastern Montpellier Snake

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Haan in Boehme 1999 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 [as M. monspessulanus] Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 Family Typhlopidae (Typical Blind Snakes)
Genus Xerotyphlops (Desert Blind Snakes)
 
Xerotyphlops vermicularis

European Worm Snake

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Dimovski 1959 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1993 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 Family Viperidae (Vipers)
Genus Vipera (Typical Vipers or Adders)
 
Vipera ammodytes

European Nose-horned Viper

Brelih & Dzukic 1974 Dimovski 1959 Geniez 2018 Heckes, Gruber & Stumpel in Joger & Stumpel 2005 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Tomovic & Dzukic 2003 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Ursenbacher, Schweiger, Tomovic, Crnobrnja-Isailovic, Fumagalli & Mayer 2008
 
Vipera berus

Northern Eurasian Viper

Brelih & Dzukic 1974 David & Ineich 1999 Dimovski 1964 Geniez 2018 Nilson, Andren & Voelkl in Joger & Stumpel 2005 Pozzi 1966 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 
Vipera ursinii

Meadow and Karst Vipers

Arnold & Ovenden 2002 Brelih & Dzukic 1974 David & Ineich 1999 Dely & Joger in Joger & Stumpel 2005 Dimovski 1964 Geniez 2018 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Nilson & Andren 2001 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski 2006 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Venchi & Sindaco 2007

Turtles (Testudines)
= endemicEX = extinct or assumed extinct

Species
References
 Family Emydidae (Western Pond Turtles)
Genus Emys (European Pond Turtles)
 
Emys orbicularis

Common European Pond Turtle

Brelih & Dzukic 1974 Fritz in Fritz (ed.) 2001 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 Family Geoemydidae (Eastern Pond Turtles)
Genus Mauremys (Stripe-necked Turtles)
 
Mauremys rivulata

Balkan Pond Turtle

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Mantziou & Rifai 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Wischuf & Busack in Fritz (ed.) 2001
 Family Testudinidae (Tortoises)
Genus Testudo (Palearctic Tortoises)
 
Testudo graeca

Spur-thighed Tortoise

Arnold & Ovenden 2002 Brelih & Dzukic 1974 Buskirk, Keller & Andreu in Fritz (ed.) 2001 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016
 
Testudo hermanni

Greek Tortoise

Bertolero, Cheylan, Hailey, Livoreil & Willemsen 2011 Brelih & Dzukic 1974 Cheylan in Fritz (ed.) 2001 Ljubisavljevic, Dzukic, Vukov & Kalezic 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016