Reptiles of KwaZulu-Natal [South Africa] Related checklists:  South Africa Bibliography of the reptiles of KwaZulu-Natal Taxonomic history of the reptiles of KwaZulu-Natal Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Monopeltis (Spade-snouted Worm Lizards)   Monopeltis sphenorhynchus Slender Spade-snouted Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley, Gans & Visser 1976 Gans 2005 Genus Zygaspis (Purple Round-headed Worm Lizards)   Zygaspis arenicola Maputo Round-headed Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 [as violacea] Bourquin 2004 Branch 1998 Broadley & Broadley 1997

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus niloticus Nile Crocodile Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthocercus (Ridgeback Agamas)   Acanthocercus atricollis Black-necked Ridgeback Agama Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Johnson & Raw 1989 Wagner, Butler, Ceriaco & Bauer 2021 Wagner, Greenbaum & Bauer 2012 Wagner, Greenbaum, Bauer, Kusamba & Leache 2018 Genus Agama (Typical Agamas)   Agama aculeata Kalahari Spiny Agama Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004   Agama atra Southern Rock Agama Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998  Family Chamaeleonidae (Chameleons) Genus Bradypodion (Southern African Dwarf Chameleons)   * Bradypodion caeruleogula Eshowe Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Glaw 2015 Raw & Brothers 2008 Tilbury 2010   Bradypodion dracomontanum Drakensberg Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010   Bradypodion melanocephalum Black-headed Dwarf Chameleon Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Glaw 2015 Johnson & Raw 1989 Klaver & Boehme 1997 Tilbury 2010   * Bradypodion nemorale Qudeni Dwarf Chameleon Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010   * Bradypodion ngomeense Ngome Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Glaw 2015 Tilbury & Tolley 2009 Tilbury 2010   Bradypodion setaroi Saint Lucia Dwarf Chameleon Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010   * Bradypodion thamnobates Natal Midlands Dwarf Chameleon Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010 Genus Chamaeleo (Typical Chameleons)   Chamaeleo dilepis Flap-necked Chameleon Bourquin 1989 Bourquin 2004 Branch 1998 Johnson & Raw 1989 Klaver & Boehme 1997 Tilbury 2010  Family Cordylidae (Girdled Lizards) Genus Chamaesaura (African Grass Lizards)   Chamaesaura aenea Coppery Grass Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998   Chamaesaura anguina Southern Highland Grass Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998   Chamaesaura macrolepis Large-scaled Grass Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Genus Cordylus (Girdled Lizards)   Cordylus cordylus Cape Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1981 Diedericks & Daniels 2014   Cordylus jonesii Limpopo Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Branch 2002   Cordylus vittifer Transvaal Girdled Lizard Bates & Broadley 2012 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Genus Platysaurus (Flat Lizards)   Platysaurus intermedius Common Flat Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1978   Platysaurus lebomboensis Lebombo Flat Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998 Jacobsen 1994 Genus Pseudocordylus (Eastern Crag Lizards)   Pseudocordylus langi Lesotho Crag Lizard Bates 2013 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998   Pseudocordylus melanotus Common Crag Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998   Pseudocordylus spinosus Spiny Crag Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998   Pseudocordylus subviridis Drakensberg Crag Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Genus Smaug (Dragon Girdled Lizards)   Smaug swazicus Swazi Girdled Lizard Bates & Stanley 2020 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as barbertonensis] Bourquin 1989 [as barbertonensis] Bourquin 2004 [as barbertonensis]   Smaug warreni Lebombo Girdled Lizard Bates & Stanley 2020 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Stanley & Bates 2014  Family Gekkonidae (Typical Geckos) Genus Afroedura (Flat Geckos)   Afroedura marleyi Zulu Flat Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Jacobsen, Kuhn, Jackman & Bauer 2014   Afroedura nivaria Drakensberg Flat Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Jacobsen, Kuhn, Jackman & Bauer 2014   Afroedura pondolia Pondo Flat Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Jacobsen, Kuhn, Jackman & Bauer 2014 Johnson & Raw 1989   * Afroedura pongola Pongola Flat Gecko Jacobsen, Kuhn, Jackman & Bauer 2014 Genus Chondrodactylus (Giant Ground Gecko and Thick-toed Geckos)   Chondrodactylus turneri Zambezi Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Heinz, Brennan, Jackman & Bauer 2021 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus mabouia Cosmopolitan House Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1977 Johnson & Raw 1989 Loveridge 1957 Maritz & Alexander 2007 Genus Homopholis (African Velvet Geckos)   Homopholis wahlbergii Southern African Velvet Gecko Bourquin 1989 Bourquin 2004 Bourquin 2019 Branch 1998 Broadley, Jackman & Bauer 2014 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus capensis Cape Dwarf Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley & Howell 1991 Johnson & Raw 1989 Loveridge 1957 Rebelo, Bates, Burger, Branch & Conradie 2019   Lygodactylus ocellatus Common Ocellated Dwarf Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Pachydactylus (Typical Thick-toed Geckos)   Pachydactylus maculatus Spotted Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998   Pachydactylus vansoni Soutpansberg Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2002 Bauer & Lamb 2005 Bourquin 1989 Bourquin 2004 Branch 1998  Family Gerrhosauridae (Plated Lizards) Genus Broadleysaurus (Rough-scaled Plated Lizard)   Broadleysaurus major Rough-scaled Plated Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1987 Loveridge 1957 Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Bourquin 1989 Bourquin 2004 Johnson & Raw 1989 Maritz & Alexander 2007 Wermuth 1968 Genus Matobosaurus (Giant Plated Lizards)   Matobosaurus validus Eastern Giant Plated Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Bourquin 1989 Bourquin 2004 Branch 1998 Genus Tetradactylus (Plated Snake Lizards)   Tetradactylus africanus African Plated Snake Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Mason & Alexander 1996   Tetradactylus breyeri Transvaal Plated Snake Lizard Bates 1996 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998   Tetradactylus seps Short-legged Plated Snake Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1981 Branch 1998  Family Lacertidae (Typical Lizards) Genus Ichnotropis (Rough-scaled Lizards)   Ichnotropis capensis Southern Rough-scaled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Genus Meroles (Southwest African Desert Lizards)   Meroles squamulosus Rough-scaled Desert Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Loveridge 1957 Genus Nucras (Sandveld Lizards)   Nucras holubi Southern Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998   Nucras intertexta Spotted Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998   Nucras lalandii Southeastern Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1981 Branch 1998   Nucras ornata Ornate Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1972 Genus Pedioplanis (African Sand Lizards)   Pedioplanis burchelli Southern Sand Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Genus Tropidosaura (South African Mountain Lizards)   Tropidosaura cottrelli Drakensberg Mountain Lizard Bates 2013 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin & Channing 1980 Bourquin 1989 Bourquin 2004 Branch 1998   Tropidosaura essexi Lesotho Mountain Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bischoff 1992 Bourquin 1989 Bourquin 2004 Branch 1998   Tropidosaura montana Common South African Mountain Lizard Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bischoff 1992 Bourquin 1989 Bourquin 2004 Branch 1998  Family Scincidae (Skinks) Genus Acontias (Greater Legless Skinks)   Acontias gracilicauda Thin-tailed Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004   Acontias parietalis Maputaland Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 [as a. aurantiacus] Bourquin 2004 [as a. aurantiacus] Branch 1998 [as a. aurantiacus] Broadley 1968 [as A. a. aurantiacus] Broadley 1990 Pietersen, Scholtz & Bastos 2018   Acontias plumbeus Giant Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as A. poecilus] Bourquin & Lambiris 1996 [as A. poecilus] Bourquin 1989 Bourquin 2004 [as A. poecilus] Branch 1998 [as A. poecilus] Broadley & Greer 1969 Johnson & Raw 1989 Lamb, Biswas & Bauer 2010 [as A. poecilus] Zhao, Verdu-Ricoy, Mohlakoana, Jordaan, Conradie & Heideman 2019   Acontias wakkerstroomensis Wakkerstroom Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as breviceps] Conradie, Busschau & Edwards 2018 Lamb, Biswas & Bauer 2010 [as breviceps] Genus Cryptoblepharus (Indo-Pacific Snake-eyed Skinks)   Cryptoblepharus africanus African Coral-rag Snake-eyed Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 [as boutonii] Bourquin 2004 Branch 1998 Haacke 1975 Genus Mochlus (African Supple Skinks)   Mochlus sundevallii Common Savanna Supple Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Loveridge 1957 Genus Panaspis (African Snake-eyed Skinks)   Panaspis wahlbergii Savanna Snake-eyed Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Jacobsen & Broadley 2000 Johnson & Raw 1989 Kilunda, Conradie, Wasonga, Jin, Peng, Murphy, Malonza & Che 2019 Maritz & Alexander 2007 Medina, Bauer, Branch, Schmitz, Conradie, Nagy, Hibbitts, Ernst, Portik, Nielsen, Colston, Kusamba, Behangana, Roedel & Greenbaum 2016 Genus Scelotes (Dwarf Burrowing Skinks)   Scelotes arenicola Zululand Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1994   Scelotes bidigittatus Lowveld Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1994   * Scelotes farquharsoni Qudeni Dwarf Burrowing Skink Raw 2020   Scelotes fitzsimonsi Kosi Bay Dwarf Burrowing Skink Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Broadley 1994   * Scelotes guentheri Natal Dwarf Burrowing Skink Armstrong 2001 [as bourquini] Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as bourquini] Bourquin 2004 [as bourquini] Branch 1998 [as bourquini] Broadley 1994 [as bourquini]   * Scelotes inornatus Plain Dwarf Burrowing Skink Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1994   Scelotes mirus Montane Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1994   Scelotes mossambicus Mozambique Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 [as brevipes] Bourquin 2004 Branch 1998 Broadley 1994 Johnson & Raw 1989 [as brevipes]   Scelotes vestigifer Coastal Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998 Broadley 1994 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis capensis Cape Skink Bourquin 1989 Bourquin 2004   Trachylepis depressa Eastern Coastal Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 2000   Trachylepis homalocephala Red-sided Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004   Trachylepis margaritifer African Rainbow Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Bauer 1998 Broadley 2000   Trachylepis punctatissima Montane Speckled Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1977   Trachylepis striata Eastern Striped Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1977 Broadley 2000 Johnson & Raw 1989 Maritz & Alexander 2007   Trachylepis varia Savanna Variable Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Johnson & Raw 1989 Maritz & Alexander 2007 Weinell & Bauer 2018  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bourquin 1989 Bourquin 2004   Varanus niloticus Nile Monitor Bourquin 1989 Bourquin 2004 Charlton 1973 Johnson & Raw 1989

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hotamboeia Common Herald Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1983 Johnson & Raw 1989 Maritz & Alexander 2007 Genus Dasypeltis (African Egg Eaters)   Dasypeltis inornata Southern Brown Egg Eater Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Gans 1959 Johnson & Raw 1989   Dasypeltis medici East African Egg Eater Bates & Broadley 2018 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Howell 1991 Broadley 1983   Dasypeltis scabra Common African Egg Eater Bourquin 1989 Bourquin 2004 Broadley 1983 Gans 1959 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa aulica Cross-barred Nocturnal Tree Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Rasmussen 1989 Genus Dispholidus (Boomslangs)   Dispholidus viridis Green Boomslang Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as D. typus] Genus Meizodon (African Smooth Snakes)   Meizodon semiornatus Semi-ornate Smooth Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Schatti 1985 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Natriciteres sylvatica Eastern Forest Marsh Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Spawls, Howell, Drewes & Ashe 2002 Genus Philothamnus (African Green Snakes)   Philothamnus angolensis Angolan Green Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983   Philothamnus hoplogaster Southeastern Green Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Johnson & Raw 1989 Maritz & Alexander 2007   Philothamnus natalensis Eastern Natal Green Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Johnson & Raw 1989   Philothamnus occidentalis Western Natal Green Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1966 [as P. natalensis occidentalis] Broadley 1983   Philothamnus semivariegatus Variegated Bush Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1983 Johnson & Raw 1989 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus semiannulatus Southern Tiger Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Genus Thelotornis (African Vine Snakes)   Thelotornis capensis Southern Vine Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Wallach 2002 Broadley 1983 Broadley 2001 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Johnson & Raw 1989 Spawls & Branch 1995  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Aspidelaps (Shield-nosed Cobras)   Aspidelaps scutatus Shield-nosed Coral Snake Reissig, Rensburg & Shaw 2015 Genus Dendroaspis (Mambas)   Dendroaspis angusticeps Southern Green Mamba Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Marais & McGahey 2010   Dendroaspis polylepis Black Mamba Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Dobiey & Vogel 2007 Haakansson & Madsen 1983 Genus Elapsoidea (African Garter Snakes)   Elapsoidea boulengeri Zambezi Garter Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993   Elapsoidea sundevallii Southern African Garter Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1971 Broadley 1983 Dobiey & Vogel 2007 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Genus Hemachatus (Ring-necked Spitting Cobra)   Hemachatus haemachates Ring-necked Spitting Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Dobiey & Vogel 2007 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Genus Naja (Typical Cobras)   Naja annulifera African Banded Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1968 Broadley 1983 Broadley 1995 Spawls & Branch 1995   Naja mossambica Mozambique Spitting Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1968 Broadley 1983 Johnson & Raw 1989 Wallach, Williams & Boundy 2014 Wuster, Crookes, Ineich, Mane, Pook, Trape & Broadley 2007   Naja subfulva Brown Forest Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as N. melanoleuca] Bourquin 1989 [as N. melanoleuca] Bourquin 2004 [as N. melanoleuca] Branch 1998 [as N. melanoleuca] Broadley 1968 [as N. melanoleuca] Broadley 1983 [as N. melanoleuca] Dobiey & Vogel 2007 [as N. melanoleuca] Hitchins & Moll 1967 [as N. melanoleuca] Johnson & Raw 1989 [as N. melanoleuca] Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis platurus Yellow-bellied Sea Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983  Family Lamprophiidae (House Snakes, etc.) Genus Alopecion (Spotted House Snake)   Alopecion guttatum Spotted House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas bicolor Bicoloured Quill-snouted Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983   Amblyodipsas concolor Natal Purple-glossed Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1970 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1971 Broadley 1983 Haacke 1982 Johnson & Raw 1989 Maritz & Alexander 2007 Raw 1971 Wallach, Williams & Boundy 2014   Amblyodipsas microphthalma Eastern Purple-glossed Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1971 Broadley 1983 Wallach, Williams & Boundy 2014   Amblyodipsas polylepis Common Purple-glossed Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1971 Broadley 1983 Maritz & Alexander 2007 Wallach, Williams & Boundy 2014   Amblyodipsas transvaalensis Transvaal Quill-snouted Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1970 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1971 Broadley 1983 Genus Amplorhinus (Many-spotted Snake)   Amplorhinus multimaculatus Many-spotted Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus capensis Cape Centipede Eater Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1983 Johnson & Raw 1989 Maritz & Alexander 2007 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis bibronii Southern Stiletto Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Broadley 1991 Maritz & Alexander 2007 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon capensis Brown House Snake Bourquin 1989 [as fuliginosus] Bourquin 2004 Broadley 1983 [as fuliginosus] Johnson & Raw 1989 [as f. fuliginosus] Loveridge 1957 [as f. fuliginosus] Maritz & Alexander 2007 [as fuliginosus] Genus Duberria (African Slug Eaters)   Duberria lutrix Common Slug Eater Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Johnson & Raw 1989 Maritz & Alexander 2007   Duberria variegata Spotted Slug Eater Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Genus Gracililima (Black File Snake)   Gracililima nyassae Black File Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1970 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Johnson & Raw 1989 Lanza & Broadley 2014 Maritz & Alexander 2007 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis nototaenia Southeastern Bark Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998 Broadley & Hughes 2000 Haagner 1990 Genus Homoroselaps (African Harlequin Snakes)   Homoroselaps dorsalis Striped Harlequin Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983   Homoroselaps lacteus Spotted Harlequin Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1983 Genus Inyoka (Swazi Rock Snake)   Inyoka swazicus Swazi Rock Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Schmidt & Scott 2003 Genus Lamprophis (SubSaharan House Snakes)   Lamprophis aurora Aurora House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983   Lamprophis fuscus Yellow-bellied House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1983 Genus Limaformosa (Venus File Snakes)   Limaformosa capensis Cape File Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum 2018 Genus Lycodonomorphus (Southern African Water Snakes)   Lycodonomorphus inornatus Olive-brown House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983   Lycodonomorphus laevissimus Dusky-bellied Water Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Haagner & Branch 1994   Lycodonomorphus obscuriventris Floodplain Water Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1983 Kyle, Alexander & Preez 2021   Lycodonomorphus rufulus Common Brown Water Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1983 Johnson & Raw 1989 Maritz & Alexander 2007 Genus Lycophidion (African Wolf Snakes)   Lycophidion capense Cape Wolf Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1976 Broadley 1983 Broadley 1996 Johnson & Raw 1989 Laurent 1968 Loveridge 1957   Lycophidion pygmaeum Pygmy Wolf Snake Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 [as L. semiannule] Bourquin 2004 Branch 1998 Broadley 1983 [as L. semiannule] Broadley 1996   Lycophidion variegatum Variegated Wolf Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1976 Branch 1998 Broadley 1983 Broadley 1996 Raw 1975 Genus Macrelaps (Natal Black Snake)   Macrelaps microlepidotus Natal Black Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Johnson & Raw 1989 Genus Montaspis (Cream-spotted Mountain Snake)   * Montaspis gilvomaculata Cream-spotted Mountain Snake Armstrong 2001 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1991 Bourquin 2004 Branch 1998 Genus Prosymna (Shovel-snout Snakes)   Prosymna bivittata Two-striped Shovel-snout Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Broadley 1980 Broadley 1983 Warner & Combrink 2010   Prosymna janii Mozambique Shovel-snout Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1970 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1980 Broadley 1983   Prosymna lineata Lineolate Shovel-snout Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1980 Broadley 1983 Huyssteen & Jordaan 2021   Prosymna stuhlmannii East African Shovel-snout Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1970 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1980 Broadley 1983 Raw 1971 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis brevirostris Short-snouted Grass Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1983 Broadley 2002 Johnson & Raw 1989 Maritz & Alexander 2007   Psammophis crucifer Cross-marked Grass Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1977 Broadley 1983 Broadley 2002   Psammophis mossambicus East African Olive Grass Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998 Broadley 1983 [as P. phillipsii] Broadley 2002 Maritz & Alexander 2007 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis subtaeniatus Western Stripe-bellied Sand Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998 Genus Psammophylax (Skaapstekers)   Psammophylax rhombeatus Spotted Skaapsteker Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1976 Broadley 1983 Genus Pseudaspis (African Mole Snake)   Pseudaspis cana African Mole Snake Bourquin 1989 Bourquin 2004 Broadley 1983 Loveridge 1957  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops conjunctus Cape Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Bourquin 1989 Bourquin 2004 Broadley & Broadley 1999 Broadley & Wallach 1997 Broadley & Watson 1976 Broadley 1983 Hahn 1980 McDiarmid, Campbell & Toure 1999   Leptotyphlops distanti Transvaal Thread Snake Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Broadley 1999 Broadley & Watson 1976 Broadley 1983 McDiarmid, Campbell & Toure 1999   Leptotyphlops incognitus Incognito Thread Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley & Broadley 1999 Broadley & Wallach 1997 Broadley & Watson 1976 Broadley 1983 Hahn 1980   Leptotyphlops scutifrons Zambezi Thread Snake Bourquin 1989 Bourquin 2004 Broadley & Broadley 1999 Broadley & Wallach 1997 Broadley & Watson 1976 Broadley 1983 Hahn 1980 Johnson & Raw 1989   Leptotyphlops sylvicolus Southern Forest Thread Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 2004 Branch 1998 Broadley & Broadley 1999 Broadley & Wallach 1997  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python natalensis Southern Rock Python Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Broadley 1984 Johnson & Raw 1989 Spawls & Branch 1995  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops bibronii Southern African Blind Snake Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Wallach 2009 Broadley 1983 Raw 1975 Wallach, Williams & Boundy 2014   Afrotyphlops fornasinii East African Blind Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Wallach 2009 Broadley 1983 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014   Afrotyphlops schlegelii Variable Giant Blind Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1970 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Wallach 2009 Broadley 1983 Hahn 1980 Genus Rhinotyphlops (African Beaked Blind Snakes)   Rhinotyphlops lalandei Cape Blind Snake Bourquin 1989 Bourquin 2004 Raw 1975 Warner & Combrink 2010  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Dobiey & Vogel 2007   Bitis atropos Cape Mountain Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 McDiarmid, Campbell & Toure 1999   Bitis gabonica Gaboon Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Dobiey & Vogel 2007 Pitman 1974 Genus Causus (Night Adders)   Causus defilippii Snouted Night Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Johnson & Raw 1989   Causus rhombeatus Rhombic Night Adder Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1983 Johnson & Raw 1989 Maritz & Alexander 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1970 Bourquin 1989 Bourquin 2004 Hughes 1975 Hughes in Bjorndal (ed.) 1982 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1970 Bourquin 1989 Bourquin 2004 Hughes in Bjorndal (ed.) 1982 Pritchard 1979  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa galeata South African Helmeted Turtle Bourquin 1989 [as subrufa] Bourquin 2004 [as subrufa] Branch 1998 [as subrufa] Iverson 1992 [as subrufa] Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Genus Pelusios (African Mud Turtles)   Pelusios castanoides Yellow-bellied Mud Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 McCord, Joseph-Ouni & Bour 2004   Pelusios rhodesianus Zambian Mud Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Boycott 2008 McCord, Joseph-Ouni & Bour 2004   Pelusios sinuatus East African Serrated Mud Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley & Boycott 2009 McCord, Joseph-Ouni & Bour 2004   Pelusios subniger East African Black Mud Turtle Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014  Family Testudinidae (Tortoises) Genus Kinixys (Hinge-back Tortoises)   Kinixys natalensis Natal Hinge-back Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Branch 1998 Broadley 1993 Iverson 1992   Kinixys spekii Eastern Savanna Hinge-back Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 Bourquin 2004 Broadley 1993 Hailey, Broadley, Boycott & Spawls 2021   Kinixys zombensis Southeastern Hinge-back Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bourquin 1989 [as belliana] Bourquin 2004 [as belliana] Branch 1998 [as belliana] Broadley 1993 [as belliana] Fritz & Havas 2006 [as belliana] Iverson 1992 McCord, Joseph-Ouni & Tabaka 2005 Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise Bourquin 1989 Bourquin 2004 Fritz & Havas 2006 Johnson & Raw 1989